For hearing-impaired university student Yuki Itose, silence has been a natural part of life since birth. Her world is small and isolated; she commutes to campus, interacts with her best friend Rin Fujishiro, and communicates through writing and text messages—a lifestyle that offers little to no change. One day, during her commute, Yuki meets fellow student Itsuomi Nagi, a multilingual travel enthusiast and friend of Rin. When Itsuomi learns of Yuki's condition, he takes it in stride, moving Yuki's heart. From this one simple gesture, Yuki and Itsuomi's lives start changing day by day as they let each other into their own worlds.

On the planet Noman's Land, reporters Meryl Stryfe and Roberto De Niro traverse the desert in search of the infamous outlaw Vash the Stampede. But the man they find near the desolate town of Jeneora Rock is a far cry from the lethal terrorist they expect. In reality, Vash is a passive and carefree drifter; he is a proponent of peace, beloved by the residents of the town. His inaccurate reputation actually stems from the widespread atrocities committed by his twin brother Millions Knives. Still, Vash is dubbed "The Humanoid Typhoon'' due to the tendency for violent chaos to follow in his wake.

Chaos soon arrives in the form of bounty hunters seeking the high price on Vash's head, and their violent pursuit poses great danger to the town and its precious power plant. Thanks to his gunslinging prowess, Vash is able to resist most of these nefarious forces. Yet he must soon face off against the ultimate evil: the unstoppable destructive power of his malevolent brother.

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Famous racehorses that have left behind worthy legacies, unique as they can be, are reincarnated as horse girls in a parallel world. In this life, they start their journey anew as they continue to race and perhaps relive the success they once lived through.

Aspiring to become the best racehorse in Japan, a horse girl named Special Week moves to Tokyo to enroll in the Tracen Academy—an institution that nurtures horse girls like her to become better racers. There, Special Week witnesses the sophisticated running style of Silence Suzuka and is inspired to become a racer like her. Shortly after, Special Week finds herself recruited into Silence Suzuka's team, Spica. From there, she begins her path to the top—one lap at a time.