Publications

My google scholar page is likely to be more up-to-date than this one. If you need help getting a pdf, please check my ResearchGate profile or just email me.

70. Tamayo, M.N., V.B. Amoroso, F.P. Coritico, D.S. Penneys, J.R.C. Callado, J.D. Horner, and P.W. Fritsch. 2024. Additions to Philippine blueberries: Three new species of Vaccinium L. (Ericaceae) from Mindanao Island. Phytotaxa 647: 19–33. [https://doi.org/10.11646/phytotaxa.647.1.2]

69. Zuntini et al. 2024. Phylogenomics and the rise of the angiosperms. Nature. [https://doi.org/10.1038/s41586-024-07324-0]

68. Penneys, D.S., F.A. Michelangeli, W.S. Judd, F. Almeda, R.D. Stone. 2023. Proposal to conserve Bellucia, nom. cons., against the additional names Myriaspora and Loreya (Melastomataceae: Henrietteeae). Taxon 72: 669–670. [https://doi.org/10.1002/tax.12969]

67. Pacifico, R., F. Almeda, D.S. Penneys, and K. Fidanza. 2022. Systematics of the Trembleya sensu stricto clade of Microlicia (Melastomataceae: Lavoisiereae). PhytoKeys 216: 1–101. [https://doi.org/10.3897/phytokeys.216.91032]

66. Ulloa Ulloa, C., F. Almeda, R. Goldenberg, G. Kadereit, F.A. Michelangeli, D.S. Penneys, R.D. Stone, and M.C. Veranso-Libalah. 2022. Melastomataceae: Global diversity, distribution, and endemism. Pp. 3–28. In: R. Goldenberg, F.A. Michelangeli, and F. Almeda (eds). Systematics, Evolution, and Ecology of Melastomataceae. Springer Nature, Cham, Switzerland. [https://doi.org/10.1007/978-3-030-99742-7_1]

65. Judd, W.S., L.C. Majure, F.A. Michelangeli, R. Goldenberg, F. Almeda, D.S. Penneys, and R.D. Stone. 2022. Morphological variability within Melastomataceae (Myrtales), including a discussion of associated terminology. Pp. 45–85. In: R. Goldenberg, F.A. Michelangeli, and F. Almeda (eds). Systematics, Evolution, and Ecology of Melastomataceae. Springer Nature, Cham, Switzerland. [https://doi.org/10.1007/978-3-030-99742-7_3]

64. Reginato, R., F. Almeda, F.A. Michelangeli, R. Goldenberg, P.W. Fritsch, R.D. Stone, and D.S. Penneys. 2022. Historical biogeography of Melastomataceae. Pp. 87–105. In: R. Goldenberg, F.A. Michelangeli, and F. Almeda (eds). Systematics, Evolution, and Ecology of Melastomataceae. Springer Nature, Cham, Switzerland. [https://doi.org/10.1007/978-3-030-99742-7_4]

63. Penneys, D.S., F. Almeda, M. Reginato, F.A. Michelangeli, R. Goldenberg, P.W. Fritsch, and R.D. Stone. 2022. A new Melastomataceae classification informed by molecular phylogenetics and morphology. Pp. 109–165. In: R. Goldenberg, F.A. Michelangeli, and F. Almeda (eds). Systematics, Evolution, and Ecology of Melastomataceae. Springer Nature, Cham, Switzerland. [https://doi.org/10.1007/978-3-030-99742-7_5]

62. Mancera, J., F. Almeda, and D.S. Penneys. 2022. Systematics of Astronieae based on morphology: Prospects for resurrecting Bamlera and incorporating Tessmannianthus. Pp. 197–217. In: R. Goldenberg, F.A. Michelangeli, and F. Almeda (eds). Systematics, Evolution, and Ecology of Melastomataceae. Springer Nature, Cham, Switzerland. [https://doi.org/10.1007/978-3-030-99742-7_8]

61. Judd, W.S., and D.S. Penneys. 2022. Systematics of tribe Henrietteeae. Pp. 219–234. In: R. Goldenberg, F.A. Michelangeli, and F. Almeda (eds). Systematics, Evolution, and Ecology of Melastomataceae. Springer Nature, Cham, Switzerland. [https://doi.org/10.1007/978-3-030-99742-7_9]

60. Michelangeli, F.A., A.N. Nicolas, G. Ocampo, R, Goldenberg, F. Almeda, W.S. Judd, E.R. Bécquer, J.D. Skean, Jr., K. Sosa, M.K. Caddah, G.M. Ionta, D.S. Penneys, M. Alvear, M. Gavrutenko, and L.C. Majure. 2022. Why recognize Miconia as the only genus in tribe Miconieae? Pp. 235–254. In: R. Goldenberg, F.A. Michelangeli, and F. Almeda (eds). Systematics, Evolution, and Ecology of Melastomataceae. Springer Nature, Cham, Switzerland. [https://doi.org/10.1007/978-3-030-99742-7_10]

59. Michelangeli, F., A.S. Dellinger, R. Goldenberg, F. Almeda, H. Mendoza-Cifuentes, D. Fernández-Fernández, C. Ulloa Ulloa, and D.S. Penneys. 2022. Phylogenetics and taxonomy of the tribe Merianieae. Pp. 255–273. In: R. Goldenberg, F.A. Michelangeli, and F. Almeda (eds). Systematics, Evolution, and Ecology of Melastomataceae. Springer Nature, Cham, Switzerland. [https://doi.org/10.1007/978-3-030-99742-7_11]

58. Penneys, D.S., and F. Almeda. 2022. An overview of the tribe Pyxidantheae. Pp. 291–305. In: R. Goldenberg, F.A. Michelangeli, and F. Almeda (eds). Systematics, Evolution, and Ecology of Melastomataceae. Springer Nature, Cham, Switzerland. [https://doi.org/10.1007/978-3-030-99742-7_13]

57. Ocampo, G., F.A. Michelangeli, D.S. Penneys, V. Handley, R. González-Moreno, E. Herrera-Dimas, and F. Almeda. 2022. A new perspective on seed morphological features in Melastomataceae. Pp. 491–531. In: R. Goldenberg, F.A. Michelangeli, and F. Almeda (eds). Systematics, Evolution, and Ecology of Melastomataceae. Springer Nature, Cham, Switzerland. [https://doi.org/10.1007/978-3-030-99742-7_23]

56. Almeda, F., and D.S. Penneys. 2022. Patterns of chromosome number diversity and evolution in the Melastomataceae. Pp. 533–561. In: R. Goldenberg, F.A. Michelangeli, and F. Almeda (eds). Systematics, Evolution, and Ecology of Melastomataceae. Springer Nature, Cham, Switzerland. [https://doi.org/10.1007/978-3-030-99742-7_24]

55. Michelangeli, F.A., A.N. Nicolas, G. Ocampo, R, Goldenberg, F. Almeda, W.S. Judd, E.R. Bécquer, J.D. Skean, Jr., R. Kriebel, K. Sosa, M.K. Caddah, G.M. Ionta, D.S. Penneys, M. Alvear, M. Gavrutenko, J.M. Burke, L.C. Majure, and M. Reginato. 2022. Historical biogeography of the Neotropical Miconieae (Melastomataceae) reveals a pattern of progressive colonization out of lowland South America. Pp. 629–644. In: R. Goldenberg, F.A. Michelangeli, and F. Almeda (eds). Systematics, Evolution, and Ecology of Melastomataceae. Springer Nature, Cham, Switzerland. [https://doi.org/10.1007/978-3-030-99742-7_28]

54. Tamayo, M.N., F.P. Coritico, V.B. Amoroso, D.S. Penneys, D.N. Tandang, and P.W. Fritsch. 2022. Vaccinium carmesinum (Ericaceae), a new blueberry species from Mt. Tago Range, Mindanao Island, Philippines. Phytotaxa 533: 173–180. [https://doi.org/10.11646/phytotaxa.533.3.3]

53. Cannon, C., C.G. Borchetta, D.L. Anderson, G. Arellano, M. Barker, G. Charron, J.M. LaMontagne, J.H. Richards, E. Abercrombie, J.E.G. Andino, L. Banin, X.T. Casapia, X. Chen, P. Degtjarenko, J. Dell, D. Durden, R. Hernandez-Gutierrez, A.D. Hirons, C.S. Kua, H. La Vigne, M. Leponce, J.Y. Lim, M.D. Lowman, A.J. Marshall, S.T. Michaletz, B.B. Normark, D.S. Penneys, G.F. Schneider, J.S. Strijk, B.B. Tiamiyu, T.L.E. Trammell, S.R. Weintraub, A.L. Desbiens, and M. Spenko. 2021. Extending our scientific reach in arboreal ecosystems for research and management. Frontiers in Forests and Global Change 4. [https://doi.org/10.3389/ffgc.2021.712165]

52. Maurin, O., A. Anest, S. Bellot, E. Biffin, G. Brewer, T. Charles-Dominique, S. Dodsworth, N. Epitawalage, B. Gallego, A. Giaretta, R. Goldenberg, D.J.P. Gonçalves, S. Graham, P. Hoch, F. Mazine, Y.W. Low, C. McGinnie, F.A. Michelangeli, S. Morris, D.S. Penneys, O.A. Pérez Escobar, Y. Pillon, L. Pokorny, G. Shimizu, V.G. Staggemeier, A.Thornhill, K.W. Tomlinson, I. Turner, T. Vasconcelos, P.G. Wilson, A.R. Zuntini, W.J. Baker, F. Forest, and Eve Lucas. 2021. A nuclear phylogenomic study of the angiosperm order Myrtales, exploring the potential and limitations of the universal Angiosperms353 probe set. American Journal of Botany 108: 1–25. [http://doi.org/10.1002/ajb2.1699]

51. Kartonegoro, A., M.C. Veranso-Libalah, G. Kadereit, A. Frenger, D.S. Penneys, S. Mota de Oliveira, and P.C. Van Welzen. 2021. Molecular phylogenetics of the Dissochaeta-alliance (Melastomataceae): Redefining tribe Dissochaeteae. Taxon 70:793–825. [https://doi.org/10.1002/tax.12508]

50. Dellinger, A.S., R. Pérez-Barrales, F.A. Michelangeli, D.S. Penneys, D. Fernández-Fernández, and J. Schönenberger. 2021. Low bee abundances explains pollinator shifts to vertebrates in tropical mountains. New Phytologist 231: 864–877. [https://doi.org/10.1111/nph.17390]

49. Galindon, J.M.M., D.S. Penneys, N.E. Lagunday, D.N. Tandang, F.P. Coritico, V.B. Amoroso, and P.W. Fritsch. 2021. Hypericum perryongii (Hypericaceae), a new species from Mindanao Island, Philippines. Phytotaxa 478: 61–66. [https://doi.org/10.11646/phytotaxa.478.1.4]

48. Fritsch, P.W., V.B. Amoroso, F. Coritico, and D.S. Penneys. 2020. Vaccinium hamiguitanense (Ericaceae), a new species from the Philippines. Journal of the Botanical Research Institute of Texas 14: 281–287. [https://doi.org/10.17348/jbrit.v14.i2.1009]

47. Michelangeli, F.A., F. Almeda, R. Goldenberg, and D.S. Penneys. 2020. A guide to curating New World Melastomataceae collections with a linear generic sequence to world-wide Melastomataceae. Preprints 2020: 2020100203. [doi:10.20944/preprints202010.0203.v1] (Not peer-reviewed)

46. Cámara-Leret, R., et al. 2020. New Guinea has the world’s richest island flora. Nature 584: 579–584. [https://www.nature.com/articles/s41586-020-2549-5]

45. Penneys, D.S., F. Almeda, F.A. Michelangeli, R. Goldenberg, A.B. Martins, and P.W. Fritsch. 2020. Lithobieae and Eriocnemeae: two new neotropical tribes of Melastomataceae. Phytotaxa 453: 157–178. [https://doi.org/10.11646/phytotaxa.453.3.1]   pdf

44. Tandang, D.N., J.M.M. Galindon, E.R. Tadiosa, F. Coritico, V. Amoroso, N.E. Lagunday, R.A.A. Bustamante, D.S. Penneys, and P.W. Fritsch. 2020. Dilochia deleoniae (Orchidaceae), a new species from Mindanao, Philippines. Phytokeys 139: 91–97. [doi:10.3897/phytokeys.139.46935]

43. Dellinger, A.S., S. Artuso, S. Pamperl, F.A. Michelangeli, D.S. Penneys, D. Fernández-Fernández, W.S. Armbruster, Y. Staedler, C. Klingenberg, and J. Schönenberger. 2019. Modularity increases rate of floral evolution and adaptive success for functionally specialized pollination systems. Communications Biology 2: 453. [doi:10.1038/s42003-019-0697-7]

42. Dellinger, A.S., L.M. Scheer, S. Artuso, D. Fernández-Fernández, F. Sornoza, D.S. Penneys, R. Tenhaken, S. Dötterl, and J. Schönenberger. 2019. Bimodal pollination systems in Andean Melastomataceae involving birds, bats, and rodents. The American Naturalist 194: 104–116. [https://doi.org/10.1086/703517]

41. Zhao, W.Y., P.W. Fritsch, V.T. Do, Q. Fan, Q.Y. Yin, D.S. Penneys, U. Swenson, and W.B. Liao. 2019. Rehderodendron truongsonense (Styracaceae), a new species from Vietnam. Journal of the Botanical Research Institute of Texas 13: 157–171. [https://www.brit.org/sites/default/files/public/brit_press/JBRIT/JBRIT-13-1-157-Zhao.pdf]

40. Michelangeli, F.A., R. Goldenberg, F. Almeda, W.S. Judd, E.R. Bécquer, G. Ocampo, G.M. Ionta, J.D. Skean, Jr., L.C. Majure, and D.S. Penneys. 2019. Nomenclatural novelties in Miconia (Melastomataceae: Miconieae). Brittonia 71: 82–121. [doi:10.1007/s12228-018-9546-0]

39. Zhang, H., C. Zuelsdorf, D.S. Penneys, S. Fan, J. Kofsky, and B.H. Song. 2018. Transcriptome profiling of a beach-adapted wild legume for dissecting novel mechanisms of salinity tolerance. Scientific Data 5: article 180290. [https://doi.org/10.1038/sdata.2018.290]

38. Almeda, F., and D.S. Penneys. 2018. Blakea echinata (Melastomataceae: Blakeeae): a new species from the Caribbean rainforest of Panama. Phytotaxa 372: 104–110. [doi: /10.11646/phytotaxa.372.1.9]

37. Almeda, F., and D.S. Penneys. 2018. A new Miconia (Melastomataceae: Miconieae) from upland rain forest of northwestern Guyana. Phytotaxa 369: 115–120. [doi: /10.11646/phytotaxa.369.2.5]

36. Dellinger, A.S., M. Chartier, D. Fernández-Fernández, D.S. Penneys, M. Alvear, F. Almeda, F.A. Michelangeli, Y. Staedler, W.S. Armbruster, and J. Schönenberger. 2018. Beyond buzz-pollination – departures from an adaptive plateau lead to new pollination syndromes. New Phytologist 221: 1136–1149. [doi.org/10.1111/nph.15468]

35. Mancera, J.P., D.S. Penneys, and F.P. Coritico. 2017. Revisiting Astrocalyx Merr. (Astronieae: Melastomataceae): New morphological observations on the Philippine endemic, enigmatic, endangered, and monotypic genus. Natural History Bulletin of the Siam Society 62: 49–65. [http://www.siam-society.org/pub_NHB/nhbss_062_1.html]

34. Fernández-Fernández, D., C. Ulloa Ulloa, and D.S. Penneys. 2016. A new species of Blakea (Blakeeae, Melastomataceae) from Ecuador. Phytotaxa 284: 69–74. [doi:10.11646/phytotaxa.284.1.7]

33. Michelangeli, F.A., F. Almeda, M. Alvear, E. Bécquer-Granados, J. Burke, M.K. Caddah, R. Goldenberg, G.M. Ionta, W.S. Judd, L.C. Majure, J. Meirelles, A.N. Nicolas, G. Ocampo, D.S. Penneys, J.D. Skean, Jr., and C. Ulloa-Ulloa. 2016. Proposal to conserve the name Miconia, nom. cons. against the additional names Maieta and Tococa (Melastomataceae, Miconieae). Taxon 65: 892–893. [doi:10.12705/654.27]

32. Almeda, F., H. Mendoza-Cifuentes, D.S. Penneys, F.A. Michelangeli, and M. Alvear. 2016. Melastomataceae. Pp. 1585–1664, 2537-2538. In: R. Bernal, S.R. Gradstein, and M. Celis (eds). Catálogo de plantas y líquenes de Colombia, Vol. II. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. Available from [2015-onwards]: http://catalogoplantasdecolombia.unal.edu.co

31. Penneys, D.S., C. Ulloa Ulloa, D.A. Neill, and D. Fernández. 2015. A new species of Chalybea (Blakeeae, Melastomataceae) from the Ecuador-Peru border. Phytotaxa 212: 264–270. [doi:10.11646/phytotaxa.212.4.2]

30. Almeda, F., H. Mendoza-Cifuentes, D.S. Penneys, F.A. Michelangeli, and M. Alvear. 2015. Melastomataceae. Pp. 3-5. In: R. Bernal, S.R. Gradstein, and M. Celis (eds). New names and new combinations for the catalogue of the plants and lichens of Colombia. Phytoneuron 22: 1–6.

29. Dellinger, A.S., D.S. Penneys, Y.M. Staedler, L. Fragner, W. Weckwerth, and J. Schoenenberger. 2014. A specialized bird pollination system with a bellows mechanism for pollen transfer and staminal food body rewards. Current Biology 24: 1615–1619. [doi:10.1016/j.cub.2014.05.056]

28. Almeda, F., and D.S. Penneys. 2014. New and reconsidered species of Tropical American Melastomataceae. Brittonia 66: 160–169. [doi:10.1007/s12228-013-9320-2]

27. Penneys, D.S. 2013. Preliminary phylogeny of the Astronieae (Melastomataceae) based on nuclear and plastid DNA sequence data, with comments on the Philippine endemic genus, Astrocalyx. Philippine Journal of Science 142 (Special Issue): 159–168.

26. Penneys, D.S., and W.S. Judd. 2013. Combined molecular and morphological phylogenetic analyses of the Blakeeae (Melastomataceae). International Journal of Plant Sciences 174: 802–817. [doi:10.1086/670011]

25. Penneys, D.S., and W.S. Judd. 2013. New combinations and a revised circumscription for the Blakeeae (Melastomataceae). PhytoKeys 20: 17–32. [doi:10.3897/phytokeys.20.4344]

24. Michelangeli, F.A., P.J.F. Guimaraes, D.S. Penneys, F. Almeda, and R. Kriebel. 2013. Phylogenetic relationships and Distribution of New World Melastomeae (Melastomataceae). Botanical Journal of the Linnean Society 171: 38–60. [doi:10.1111/j.1095-8339.2012.01295.x]

23. Renner, S.S., D. Triebel, F. Almeda, R. Goldenberg, H. Mendoza-Cifuentes, F.A. Michelangeli, D.S. Penneys, R.D. Stone, and C. Ulloa. 2012. MELnet: Melastomataceae.Net (version 1.0, May 2011). In: F. Bisby, Y. Roskov, A. Culham, T. Orrell, D. Nicolson, L. Paglinawan, N. Bailly, W. Appeltans, P. Kirk, T. Bourgoin, G. Baillargeon, D. Ouvrard (eds). Species 2000 & ITIS Catalogue of Life, 30th April 2012. Digital resource at www.catalogueoflife.org/col/. Species 2000: Reading, UK.

22. Penneys, D.S., and E. Cotton. 2011. Melastomataceae. Pp. 393–423. In: R. Valencia, N. Pitman, S. León-Yánez, and P.M. Jørgensen (eds). Libro rojo de las plantas endémicas del Ecuador, II Edition. Publicaciones del Herbario QCA, Pontifícia Universidad Católica del Ecuador, Quito.

21. Judd, W.S., G.M. Ionta, J.D. Skean, Jr., K. Campbell, and D.S. Penneys. 2011. Noteworthy collections of Heterotis rotundifolia in Jamaica and Dominica. Castanea 76: 311–312.

20. Wanntorp, L., C. Puglisi, D.S. Penneys, and L. Ronse De Craene. 2011. Multiplications of floral organs in flowers – a case study in Conostegia (Melastomataceae, Myrtales). Pp. 218–235. In: L. Wanntorp and L. Ronse De Craene (eds). Flowers on the Tree of Life. Systematics Association Special Volume, Cambridge University Press, Cambridge, England.

19. Penneys, D.S., and W.S. Judd. 2011. Phylogenetics and morphology in the Blakeeae (Melastomataceae). International Journal of Plant Sciences 172: 78–106. [doi:10.1086/657284]

18. Penneys, D.S., W.S. Judd, F.A. Michelangeli, and F. Almeda. 2010. Henrietteeae (Melastomataceae): A new Neotropical berry-fruited tribe. Systematic Botany 35: 783–800. [doi:10.1600/036364410X539862]

17. Morales-Puentes, M.E., and D.S. Penneys. 2010. New species of Chalybea Naud. and Huilaea Wurdack. Brittonia 62: 26–34. [doi:10.1007/s12228-009-9092-x]

16. Penneys, D.S., and L. Jost. 2009. Blakea attenboroughii (Melastomataceae: Blakeeae): A new species from Ecuador. Proceedings of the California Academy of Sciences 60: 69–72.

15. Goldenberg, R., D.S. Penneys, F. Almeda, W.S. Judd, and F.A. Michelangeli. 2008. Phylogeny of Miconia (Melastomataceae): Patterns of stamen diversification in a megadiverse Neotropical genus. International Journal of Plant Science 169: 963–979. [doi:10.1086/589697]

14. Judd, W.S., J.D. Skean, Jr., D.S. Penneys, and F.A. Michelangeli. 2008. A new species of Henriettea (Melastomataceae) from the Sierra de Baoruco, the Domincan Republic. Brittonia 60: 217–227. [doi:10.1007/s12228-008-9021-4]

13. Varassin, I. G., D.S. Penneys, and F.A. Michelangeli. 2008. Comparative anatomy and morphology of nectar-producing Melastomataceae. Annals of Botany 102: 899–909. [doi:10.1093/aob/mcn180]

12. Bécquer-G., E. R., K.M. Neubig, W.S. Judd, F.A. Michelangeli, J.R. Abbott, and D.S. Penneys. 2008. Preliminary molecular phylogenetic studies in Pachyanthus (Miconieae, Melastomataceae). Botanical Review 74: 37–52. [doi:10.1007/s12229-008-9010-z]

11. Michelangeli F.A., W.S. Judd, D.S. Penneys, J.D. Skean, Jr., E.R. Bécquer-G., R. Goldenberg, and C.V. Martin. 2008. Multiple events of dispersal and radiation of the tribe Miconieae (Melastomataceae) in the Caribbean. Botanical Review 74: 53–77. [doi:10.1007/s12229-008-9004-x]

10. Almeda, F., P.E. Berry, A. Freire-Fierro, A. Gröger, B.K. Holst, N.G. Luckana, F.A. Michelangeli, T. Morley, D.S. Penneys, S.S. Renner, O.R. Robinson, J.J. Wurdack, and K. Yatskievych. 2007. Melastomataceae. Pp. 397–417. In: V. Funk, T. Hollowell, P. Berry, C. Kelloff, and S.N. Alexander (eds.). Checklist of the plants of the Guiana Shield (Venezuela: Amazonas, Bolivar, Delta Amacuro; Guyana, Surinam, French Guiana). Contributions from the United States National Herbarium, Washington D.C.

9. Blanco, M.A., W.M. Whitten, D.S. Penneys, N.H. Williams, K.M. Neubig, and L. Endara. 2006. A simple and safe method for rapid drying of plant specimens using forced-air space heaters. Selbyana 27: 83–87.

8. Penneys, D.S., and W.S. Judd. 2005. A cladistic analysis and systematic revision of Charianthus (Miconieae: Melastomataceae) using morphological and molecular characters. Systematic Botany 30: 559–584. [doi:10.1600/0363644054782125]

7. Judd, W.S., and D.S. Penneys. 2004. Taxonomic studies in the Miconieae (Melastomataceae). VIII. A revision of the species of the Miconia desportesii complex on Hispaniola. Rhodora 106: 124–147.

6. Michelangeli, F.A., D.S. Penneys, J. Giza, D.E. Soltis, M.H. Hils, and J.D. Skean, Jr. 2004. A preliminary phylogeny of the tribe Miconieae (Melastomataceae) based on nrITS sequence data and its implications on inflorescence position. Taxon 53: 279–290.

5. Penneys, D.S., and W.S. Judd. 2004. Two new species of Charianthus (Melastomataceae: Miconieae) from the Lesser Antilles. Brittonia 56: 19–27. [doi:10.1663/0007-196X(2004)056[0151:TNSOCM]2.0.CO;2]

4. Judd, W.S., D.S. Penneys, and J.D. Skean, Jr. 2004. Rediscovery of Ossaea alloeotricha, an endemic of the high-elevation Massif de la Hotte, Haiti, and its transfer to Miconia (Melastomataceae: Miconieae). Brittonia 56: 56–62. [doi:10.1663/0007-196X(2004)056[0159:ROOAAE]2.0.CO;2]

3. Penneys, D.S., and W.S. Judd. 2003. The resurrection and lectotypification of Tetrazygia fadyenii (Melastomataceae: Miconieae): A hummingbird-pollinated treelet endemic to Jamaica. Sida 20: 877–884.

2. Penneys, D.S. 2001 - 2022. Melastomataceae of the World [online]. Available at: http://www.melastomes.com (not peer-reviewed)

1. Penneys, D.S. 1996. A guide to the common flowering plant families of Monteverde, Costa Rica. Published by the author. 65 pp. (not peer-reviewed)