Speech, Language and Cognition in
Brains, Minds and Machines



Krauss, P.
Künstliche Intelligenz und Hirnforschung - Neuronale Netze, Deep Learning und die Zukunft der Kognition
Springer 2023

Peer-reviewed Journal Articles and Conference Papers

Under Review

Schilling, A., Gerum, R., Boehm, C., Rasheed, J., Metzner, C., Maier, A., Reindl, C., Hamer, H., & Krauss, P.
Deep learning based decoding of local field potentials.
preprint: doi.org/10.1101/2022.10.14.512209 

Metzner, C., Schilling, A., & Krauss, P.
Beyond Labels: Advancing Cluster Analysis with the Entropy of Distance Distribution (EDD).
preprint: arxiv.org/abs/2311.16621 

Stoewer, P., Schilling, A., Maier, A., & Krauss, P.

Multi-Modal Cognitive Maps based on Neural Networks trained on Successor Representations.

preprint: arxiv.org/abs/2401.01364 

Tziridis, K., Rasheed, J., Krauss, P., Schilling, A., & Schulze, H.
Tinnitus is associated with increased extracellular matrix density in the auditory cortex.
preprint: essopenarchive.org/doi/full/10.22541/au.170020631.15815554 


Metzner, C., Yamakou, M., Voelkl, D., Schilling, A., & Krauss, P. (2024)
Quantifying and maximizing the information flux in recurrent neural networks.
Neural Computation

Schüller, A., Schilling, A., Krauss, P., & Reichenbach, T. (2024)
The Early Subcortical Response at the Fundamental Frequency of Speech Is Temporally Separated from Later Cortical Contributions.
Journal of Cognitive Neuroscience.



Surendra, K., Schilling, A., Stoewer, P. Maier, A., & Krauss, P. (2023)
Word class representations spontaneously emerge in a deep neural network trained on next word prediction.
Proceedings of the 2023 IEEE International Conference on Machine Learning and Applications (ICMLA 2023)

Stoewer, P., Schilling, A., Maier, A., & Krauss, P. (2023)
Conceptual Cognitive Maps Formation with Neural Successor Networks and Word Embeddings.
Proceedings of the 2023 IEEE International Conference on Development and Learning (ICDL 2023)

Schilling, A., Sedley, W., Gerum, R., Metzner, C., Tziridis, K., Maier, A., Schulze, H., Zeng, F.-G., Friston, K.J., & Krauss, P. (2023)
Predictive coding and stochastic resonance as fundamental principles of auditory phantom perception.

Gerum, R., Erpenbeck, A., Krauss, P., & Schilling, A. (2023).
Leaky-Integrate-and-Fire Neuron-Like Long-Short-Term-Memory Units as Model System in Computational Biology.
2023 International Joint Conference on Neural Networks (IJCNN 2023), IEEE
awarded with the Best Paper Award among more than 1000 accepted papers

Koelbl, N., Schilling, A., & Krauss, P. (2023)
Adaptive ICA for Speech EEG Artifact Removal.
BioSMART 2023, 5th International Conference on Bio-engineering for Smart Technologies, IEEE
awarded with the Best Student Paper Award

Schilling, A., Schaette, R., Sedley, W., Gerum, R., Maier, A., & Krauss, P. (2023)
Editorial: Auditory Perception and Phantom Perception in Brains, Minds and Machines.
Frontiers in Neuroscience

Metzner, C., Schilling, A., Traxdorf, M., Schulze, H., Tziridis, K., & Krauss, P. (2023)
Extracting continuous sleep depth from EEG data without machine learning.
Neurobiology of Sleep and Circadian Rhythms.

Stoewer, P., Schilling, A., Maier, A., & Krauss, P. (2023).
Neural Network based Formation of Cognitive Maps of Semantic Spaces and the Putative Emergence of Abstract Concepts.
Scientific Reports

Schilling, A., Tziridis, K., Schulze, H., & Krauss, P. (2023).
Behavioral assessment of Zwicker tone percepts in rodents.

Stoll, A., Krauss, P., Maier, A., Gerum, R., & Schilling, A. (2023).
Coincidence Detection and Integration Behavior in Spiking Neural Networks.
Cognitive Neurodynamics

Schulze, H., Schilling, A., Krauss, P. & Tziridis,K. (2023)
The Erlangen model of tinnitus development—New perspective and treatment strategy.

Schüller, A., Schilling, A., Krauss, P., Rampp, S., & Reichenbach, T. (2023)
Attentional modulation of the cortical contribution to the frequency-following response evoked by continuous speech.
Journal of Neuroscience


Metzner, C., Schilling, A., Traxdorf, M., Tziridis, K., Schulze, H. & Krauss, P. (2022).
Classification at the Accuracy Limit -- Facing the Problem of Data Ambiguity.
Scientific Reports

Garibyan, A., Schilling, A., Boehm, C., Zankl, A., & Krauss, P. (2022).
Neural Correlates of Linguistic Collocations During Continuous Speech Perception.
Frontiers in Psychology

Schillng, A. & Krauss, P. (2022).
Tinnitus is associated with improved cognitive performance and speech perception -- Can stochastic resonance explain?
Frontiers in Aging Neuroscience

Stoewer, P., Schlieker, C., Schilling, A., Metzner, C., Maier, A., & Krauss, P. (2022).
Neural Network based Successor Representations to form Cognitive Maps of Space and Language.
Scientific Reports

Schilling, A., Gerum, R., Zankl, A., Metzner, C., Maier. A., & Krauss, P. (2022).
Intrinsic noise improves speech recognition in a computational model of the auditory pathway.
Frontiers in Neuroscience

Metzner, C. & Krauss, P. (2022).
Dynamics and Information Import in Recurrent Neural Networks.
Frontiers in Computational Neuroscience

Maier, A., Köstler, H., Heisig, M., Krauss, P. & Yang, S.H. (2022).
Known Operator Learning and Hybrid Machine Learning in Medical Imaging -- A Review of the Past, the Present, and the Future

Progress in Biomedical Engineering


Tziridis, K., Brunner, S., Schilling, A., Krauss, P. & Schulze, H. (2022).
Spectrally Matched Near-Threshold Noise for Subjective Tinnitus Loudness Attenuation Based on Stochastic Resonance.
Frontiers in Neuroscience


Boensel, F., Krauss, P., Metzner, C. & Yamakou, M. (2022).
Control of noise-induced coherent oscillations in three-neuron motifs.
Cognitive Neurodynamics


Krauss, P., Metzner, C., Joshi, N., Schulze, H., Traxdorf, M., Maier, A. & Schilling, A. (2021).

Analysis and Visualization of Sleep Stages based on Deep Neural Networks.

Neurobiology of Sleep and Circadian Rhythms


Krauss, P. and Tziridis, K. (2021).

Simulated transient hearing loss improves auditory sensitivity.
Scientific Reports


Schilling, A., Tomasello, R., Henningsen-Schomers, M.R., Zankl, A., Surendra, K., Haller, M., Karl, V., Uhrig, P., Maier, A. & Krauss, P. (2021).

Analysis of continuous neuronal activity evoked by natural speech with computational corpus linguistics methods.

Language, Cognition and Neuroscience


Metzner, C., Schilling, A., Traxdorf, M., Schulze, H. & Krauss, P. (2021).
Sleep as a random walk – Superstatistical analysis of EEG data across sleep stages.
Communications Biology

Schilling, A., Metzner, C., Gerum, R., Maier, A., & Krauss, P. (2021).

Quantifying the separability of data classes in neural networks.

Neural Networks


Yang, Z., Schilling, A., Maier, A. & Krauss, P. (2021).

Neural Networks with Fixed Binary Random Projections Improve Accuracy in Classifying Noisy Data.

Proceedings, German Workshop on Medical Image Computing, Regensburg, March 7-9, 2021


Schilling, A., Tziridis, K., Schulze, H. & Krauss, P. (2021).

The Stochastic Resonance model of auditory perception: A unified explanation of tinnitus development, Zwicker tone illusion, and residual inhibition.

Progress in Brain Research


Jeschke, M., Happel, M.F.K., Tziridis, K., Krauss, P., Schilling, A., Schulze, H. & Ohl, F.W. (2021).

Acute and long-term circuit-level effects in the auditory cortex after sound trauma.

Frontiers in Neuroscience


Tziridis, K., Forster, J., Buchheidt-Doerfler, I., Krauss, P., Schilling, A., Wendler, O., Sterna, E. & Schulze, H. (2021).
Tinnitus development is associated with synaptopathy of inner hair cells in Mongolian gerbils.
European Journal of Neuroscience


Krauss, P. & Maier, A. (2020).

Will we ever have conscious Machines?

Frontiers in Computational Neuroscience


Gerum, R., Erpenbeck, A., Krauss, P. & Schilling, A. (2020).

Sparsity through evolutionary pruning prevents neuronal networks from overfitting.

Neural Networks


Knipper, M., van Dijk, P., Schulze, H., Mazurek, B., Krauss, P., Scheper, V., Warnecke, A., Schlee, W., Schwabe, K., Singer, W., Braun, C., Delano, P.H., Fallgatter, A.J., Ehlis, A.-C., Searchfield, G.D., Munk, M.H.J., Baguley, D.M. & Rüttiger, L. (2020).

The neural bases of tinnitus: Lessons from deafness and cochlear implants

Journal of Neuroscience


Schilling, A., Krauss, P., Hannemann, R., Schulze, H. & Tziridis, K. (2020).

Reducing tinnitus intensity: Pilot study to attenuate tonal tinnitus using individually spectrally optimized near-threshold noise.




Krauss, P., Prebeck, K., Schilling, A. & Metzner, C. (2019).

'Recurrence Resonance' in three-neuron motifs.

Frontiers in Computational Neuroscience


Krauss, P., Schuster, M., Dietrich, V., Schilling, A, Schulze, H. & Metzner, C. (2019).

Weight statistics controls dynamics in recurrent neural networks.

PloS one


Krauss, P., Zankl, A., Schilling, A., Schulze, H. & Metzner, C. (2019).

Analysis of structure and dynamics in three-neuron motifs.

Frontiers in Computational Neuroscience


Krauss, P., Schilling, A., Tziridis, K. & Schulze, H. (2019).

Models of tinnitus development. From cochlea to cortex.

HNO Journal


Traxdorf, M., Krauss, P., Schilling, A., Schulze, H. & Tziridis, K. (2019).

Microstructure of cortical activity during sleep reflects respiratory events and state of daytime vigilance.



Dierich, M., Hartmann, S., Dietrich, N., Moeser, P., Brede, F., Johnson Chacko, L., Tziridis, K., Schilling, A., Krauss, P., Hessler, S., Karch, S., Schrott-Fischer, A., Blumer, M., Birchmeier, C., Oliver, D., Moser, T., Schulze, H., Alzheimer, C., Leitner, M. & Huth, T. (2019).

β-secretase BACE1 is required for normal cochlear function.

Journal of Neuroscience


Gerum, R., Rahlfs, H., Streb, M., Krauss, P., Grimm, J., Metzner, C., Tziridis, K., Günther, M., Schulze, H., Kellermann, W. & Schilling, A. (2019).

Open(G)PIAS: An open source solution for the construction of a high-precision Acoustic-Startle-Response (ASR) setup for  tinnitus screening and threshold estimation in rodents.

Frontiers in Behavioral Neuroscience


Schilling, A, Gerum, R., Krauss, P., Metzner, C., Tziridis, K. & Schulze, H. (2019).

Objective estimation of sensory thresholds based on neurophysiological parameters.

Frontiers in Neuroscience



Krauss, P., Tziridis, K., Schilling, A. & Schulze, H. (2018).

Cross-modal stochastic resonance as a universal principle to enhance sensory processing.

Frontiers in Neuroscience


Krauss, P., Schilling, A., Bauer, J., Tziridis, K., Metzner, C., Schulze, H. & Traxdorf, M. (2018).

Analysis of multichannel EEG patterns during human sleep: a novel approach.

Frontiers in Human Neuroscience


Krauss, P., Metzner, C., Schilling, A., Tziridis, K., Traxdorf M., Wollbrink, A., Rampp, S., Pantev, C., & Schulze, H. (2018).

A statistical method for analyzing and comparing spatiotemporal cortical activation patterns.

Scientific Reports


Metzner, C., Lange, J., Krauss, P., Wunderling, N., Übelacker, J., Martin, F. & Fabry, B. (2018).

Pressure-driven collective growth mechanism of planar cell colonies.

Journal of Physics D: Applied Physics



Krauss, P., Metzner, C., Schilling, A., Schütz, C., Tziridis, K., Fabry, B., & Schulze, H. (2017).

Adaptive stochastic resonance for unknown and variable input signals.

Scientific Reports


Krauss, P., Schulze, H., & Metzner, C. (2017).

A chemical reaction network to generate random, power-law distributed time intervals.

Artificial Life


Schilling, A., Krauss, P., Gerum, R., Metzner, C., Tziridis, K., & Schulze, H. (2017).

A new statistical approach for the evaluation of gap-prepulse inhibition of the acoustic startle reflex (GPIAS) for tinnitus assessment.

Frontiers in Behavioral Neuroscience


Gollnast, D., Tziridis, K., Krauss, P., Schilling, A., Hoppe, U., & Schulze, H. (2017).

Analysis of audiometric differences of patients with and without tinnitus in a large clinical database.

Frontiers in Neurology



Krauss, P., Tziridis, K., Metzner, C., Schilling, A., Hoppe, U., & Schulze, H. (2016).

Stochastic resonance controlled upregulation of internal noise after hearing loss as a putative cause of tinnitus-related neuronal hyperactivity.

Frontiers in Neuroscience


Krauss, P., Tziridis, K., Buerbank, S., Schilling, A., & Schulze, H. (2016).

Therapeutic Value of Ginkgo biloba Extract Egb 761 in an Animal Model (Meriones unguiculatus) forNoise Trauma Induced Hearing Loss and Tinnitus.

PLoS one



Lang, N. R., Münster, S., Metzner, C., Krauss, P., Schürmann, S., Lange, J., & Fabry, B. (2013).

Estimating the 3D pore size distribution of biopolymer networks from directionally biased data.

Biophysical Journal



Krauss, P., Metzner, C., Lange, J., Lang, N., & Fabry, B. (2012).

Parameter-free binarization and skeletonization of fiber networks from confocal image stacks.

PLoS one