Publications

A. Publications

Wangmo, K, & Dorji, U. (2025). Ecological Assessment of Parasites and Environmental Drivers in Native Freshwater Fishes of Punatsangchhu River, Bhutan. South Asian Journal of Parasitology, 8 (3), 255-68. https://doi.org/10.9734/sajp/2025/v8i3232. 

Masud, S.M., Gurung, D.B., & Dorji, U. (2025). Habitat Preference and Foraging Technique of River Lapwing (Vanellus duvaucelii Lesson, 1826) in Punatsangchhu River, Bhutan. Bhutan Journal of Natural Resources and Development, 12(1), 30–42. Retrieved from https://www.bjnrd.org/index.php/bjnrd/article/view/182

Dorji, U. (2025). Introduction of Brown Trout Displaces Native Fish in Bhutan – Consequences and Actions. Centre for Rural Development Studies. Designed by Kuensel. Policy brief. https://www.cnr.edu.bt/wp-content/uploads/2025/06/Brown-Trout.pdf

 Tenzin, S., & Dorji, U. .(2025). Investigating Sediment Grain Size Distribution and Transport Patterns in Punatsangchhu River, Punakha, Bhutan. Indonesian Journal of Social and Environmental Issues (IJSEI), 6(1), 94-111. https://doi.org/10.47540/ijsei.v6i1.1615

Dorji, U., & Sagar, L. (2025). Hydrological dynamics and brown trout habitat selection in the Thimphu Chhu River, Bhutan. Asian Journal of Environment & Ecology, 24(3), 51-63. https://doi.org/10.9734/ajee/2025/v24i3669

Dorji , U. (2024). Environmental DNA (eDNA) as a Tool for Freshwater Fish Biodiversity Monitoring in Bhutan: A Review. Bhutan Journal of Natural Resources and Development, 11(2), 72–87. Retrieved from https://www.bjnrd.org/index.php/bjnrd/article/view/153

Gurung, D. B., Dorji, U., Dorjee, S., (2024). Hydropower and Aquatic Ecosystems: A Study of Macroinvertebrate Diversity and River Health in Southern Bhutan. International Journal of Innovative Research in Science, Engineering and Technology 13(9), 15901-15908. https://doi.org/10.15680/IJIRSET.2024.1309001    

Wangmo, K., Sagar, L., Gurung, D. B., Dorji, U., Wangchuk, K., Dorji, R., & Subedi, R. (2024). Species Abundance and Environmental Association of Nemacheilidae in South-Central Bhutan. Bhutan Journal of Natural Resources and Development, 11(1), 34–47. Retrieved from https://www.bjnrd.org/index.php/bjnrd/article/view/134

Dorji, T., & Dorji, U. (2024). Diversity and Distribution of Bryophytes along Mochhu River Basin in Pasaphu, under Trashigang District. Bhutan Journal of Natural Resources and Development, 11(1), 1–13. Retrieved from https://www.bjnrd.org/index.php/bjnrd/article/view/120

Dorji, U., Wangchuk, K., Moktan, S., & Tenzin, U. (2023). Freshwater Metacommunity Structure of Suchhu River, Haa District, Bhutan. Bhutan Journal of Natural Resources & Development, 10(2), 28–40. https://doi.org/10.17102/cnr.2023.90

Dorji, P., Gyeltshen, P., Tobgay, K., Dorji, S., Dorji, U., Doya K.S. & Kumar, P. (2023). Additions to the orchid flora of Bhutan III: Paphiopedilum × pradhanii. Lankesteriana, 23(3), 425–432. http://dx.doi.org/10.15517/lank.v23i3.57038. 

Lhamo, S., Thinley, U., & Dorji, U. (2023). Impacts of Invasion by Ageratina adenophora on Soil Properties and Plant Diversity. Bhutan Journal of Natural Resources & Development, 10(1), 1–9. https://www.bjnrd.org/index.php/bjnrd/article/view/98/90. 

Penjor, R., Katel, O., Dorji, U., Kuman, S. & Dorji, S. (2023) Household solid waste generation and management systems in rural areas of Bhutan. The Journal of Solid Waste Technology and Management 49(2): 21179–21189. http://doi.org/10.5276/jswtm/iswmaw.    

Lhamo, S., Thinley, U., & Dorji, U. (2023). Spatio-Temporal Projection of Invasion Using Machine Learning Algorithm-MaxEnt.Journal of Forest and Environmental Science, 39(2), 105–117. https://doi.org/10.7747/JFES.2023.39.2.105.  

Wangmo, S. and Dorji, U. & Dorji, N. (2023). Assessing the Livelihood Vulnerability to Impact of Climate Change in Western Bhutan. Journal of Agriculture and Ecology Research International, 24 (6). pp. 21-39. ISSN 2394-1073. https://doi.org/10.9734/jaeri/2023/v24i6558

Dorji, P., Tashi, S., Dorji, U., Tshering, K., Tshomo, U. & Gyeltshen, T.  (2023) Diversity of Pollinator Species and Efficacy in Pollination of Strawberry yield. The Journal of Agricultural Sciences 18(3). https://doi.org/10.4038/jas.v18i3.10055   

Nima & Dorji, U. (2022). Waterbird assemblage along Punatsangchhu River, Punakha and Wangdue Phodrang, Bhutan. Journal of Threatened Taxa 14(6): 21179–21189. https://doi.org/10.11609/jott.7681.14.6.21179-21189. 

Dorji, U., & Gurung, D.B. (2022). From Conservation to Business: Urka Bangala Group, Trashi Yangtse Dzongkhag. Case Studies of Successful Farmers, Agri-enterprises, Farmers’ Groups and Cooperatives in Bhutan (pp. 62-64). College of Natural Resources: Centre for Sustainable Mountain Agriculture, ISBN: 978-99936-994-3-9.  

Dorji, U., & Gurung, D.B. (2022). Sonam Gonor Detshen, Yangtse Gewog, Trashi Yangtse Dzongkhag. Case Studies of Successful Farmers, Agri-enterprises, Farmers’ Groups and Cooperatives in Bhutan (pp. 116-119). College of Natural Resources: Centre for Sustainable Mountain Agriculture, ISBN: 978-99936-994-3-9.  

Wangmo, N., Dorji, U., & Katel, O. (2022). Climate Change and Water Sources: A Case of Phobjikha and Gangtey Gewog, Wangdue Phodrang Dzongkhag, Bhutan. Indonesian Journal of Social and Environmental Issues (IJSEI), 3(1), 3748.https://doi.org/10.47540/ijsei.v3i1.442

Tashi, S.  & Dorji, U. . (2022). Agro-virtual water trade in relation to water availability and attributes of crop import of   Lungnyi            Gewog,      Bhutan. Bhutan Journal of         Research          and       Development, 11(2). https://doi.org/10.17102/bjrd.rub.11.2.033  

Dorji, U. & Wangchuk, K. (2022). Abundance, Distribution and Association of Paphiopedilum fairrieanum (Lindl.) Stein with Site Factors in Brongshing, Samdrup Jongkhar District. Conservation, 2,627–638. https://doi.org/10.3390/conservation2040041     

Tashi, S., Gurung, D.B., Dorji, U. and Wangchuk, K. (2022). Trace elements in fish species: Schizothorax richardsonii, Salmo trutta and Neolissochilus hexagonolepis in relation to sizes and tissues of Punatsang Chhu. Environmental Chemistry and Ecotoxicology, Science Direct, Elsevier. 4(2022):78-88. https://doi.org/10.1016/j.enceco.2022.01.001. ISSN 25901826. 

Nepal, A., Tashi, S., Chhetri, R., Dorji, T., Dorji, U., & Sapkota, S. (2021). Impacts of Climate Change on Hazelnut (Corylus avellane L.) Cultivation in Bhutan. Vietnam Journal of Agricultural Sciences, 5(2). retrieved from https://vjas.vnua.edu.vn/index.php?journal=vjas&page=article&op=view&path[]=334   

Dorji, U. (2022). Abstraction of Groundwater in Mountainous Regions. Kuensel, 14 February 2022. https://kuenselonline.com/abstraction-of-groundwater-in-mountainous-regions/  

Dorji, G. and Dorji, U. (2022). Habitat use, feeding behaviour and conservation threats of Black-necked Cranes at Gangtey-Phobji Valley, Wangdue Phodrang, Bhutan. International Journal of Environment. 10 (2): 64-82. 

http://dx.doi.org/10.3126/ije.v10i1.38404. ISSN 2091-2854. 

Dorji, U. (2022). How safe is the treatment of our drinking water?  Kuensel Corporation Limited: Thimphu. ISBN 978-99936-994-0-8.  

https://kuenselonline.com/how-safe-is-the-treatment-of-our-drinking-

water/?fbclid=IwAR0Vzqqks4nzT2hwOyK32-hyM1pSt_uZiIanw_3exy7ydb9PHIFmtglCxRg

Dorji, U. (2021). Freshwater fish diversity and Water Quality assessment in Suchhu River, Haa.  Kuensel Corporation Limited: Thimphu. ISBN 978-99936-994-0-8.         https://www.researchgate.net/project/Water-Quality-and-Aquatic-EcologyAssessments-for-14-MW-Suchhu-Hydropower-Project-Haa

Dorji, U. (2021). Freshwater fish diversity and Water Quality assessment in Burgangchhu River, Haa. Druk Green Power 

Corporation            Limited:            Thimphu.  International Journal of Environment. 10 (2): 64-82. https://www.researchgate.net/project/Water-Quality-and-Aquatic-EcologyAssessments-for-48-MW-Burgangchhu-Hydropower-Project-Zhemgang.  

Ghalley, T.B, Dorji, U., Dorji, C., Nepal, A. and Shacha, N. (2021). Water Beetles: species diversity and response to physico-chemical parameters in different freshwater habitats in Samtse, Bhutan. International Journal of Environment. 10 (1): 64-82. http://dx.doi.org/10.3126/ije.v10i1.38404. ISSN 2091-2854. 

Tshering K. & Dorji, U., (2021). Habitat Assessment of Shilajit (Asphaltum punjabianum L.) in Khateo-Khame Gewog, Gasa Dzongkhag. Annual Research Meet, College of Natural Resources, Centre for Environment and Climate Research.  

Dorji, U. (2021). Water Resources Management in Bhutan. Sustainable Natural Resource Management in the Himalayan Region: Livelihood and Climate Change. Nova Science Publishers ISBN: 978-1-53618-962-9 

Thinley, J., Dorji, U.,  Tshering, U.,  Nepal , A.,  Chaida, L., Chaten.,  Rai, K., Tobgay, S., Tshering, B., Dorji, T.,  Thinley, P.,  Rabgay, T.,  Yongdrup, P. and  Wangchuk, Y. (2021) Mineral Composition and Behaviour of Mammals at Natural Saltlicks in Jomotsangkha Wildlife Sanctuary, Bhutan. Bhutan Journal of Natural Resources & Development. 7(2): 23-33. https://doi.org/10.17102/cnr.2020.49

Tenzin, U. & Dorji, U. (2020). Distributional Pattern and Environmental Response of Snail in Central Southern Bhutan. Bhutan Journal of Natural Resources & Development. 7(1): 1-11. https://doi.org/10.17102/cnr.2020.41. ISSN 2409–5273. 

Dorji, U. (2020). Aquatic ecological Impact Assessment as part of Environment and Social Impact Assessment of Gelephu Flood Protection Project. Sarpang: Gelephu Thromde.  Kuensel Corporation Limited: Thimphu. ISBN 978-99936-994-0-8.  https://www.researchgate.net/project/Aquatic-ecological-Impact-Assessment-as-part-of-Environment-and-Social-Impact-Assessment-of-Gelephu-Flood-Protection-Project  

Nepal, A. & Dorji, U. (2020). Composition and Diversity Pattern of Climbers in Tropical Forest of Langchenphu, Jomotshangkha Wildlife Sanctuary. Bhutan Journal of Natural Resources & Development. 7(1): 40-50. 

https://doi.org/10.17102/cnr.2020.44. ISSN 2409–5273. 

Dorji, U. (2019). Assessment of Fish Diversity and Water Quality Parameters for 1125 MW Dorjilung Hydropower Project. Thimphu: Druk Green Power Corporation of Bhutan. Kuensel Corporation Limited: Thimphu. ISBN 978-99936-994-0-8.  https://www.researchgate.net/project/Assessment-of-FishDiversity-and-Water-Quality-Parameters-1125-MW-Dorjilung-Hydropower-Project  

Dahal, D., & Dorji, U. (2019). Comparative Analysis of Water Discharge Measurement Methods and Seasonal Trend Analysis of Taebayrongchu Sub-watershed, Punakha. Bhutan Journal of Research & Development, 8(2), 39-54.  ISSN 1321- 4585.  https://www.rub.edu.bt/index.php/en/journals/662-bjrd-autumn-2019-vol-8-no-2.html  

Lhundup, K. & Dorji, U. (2018). Macro-invertebrate Diversity and its relationship with environmental variables in Adha Lake between monsoon and post-monsoon seasons. Bhutan Journal of Natural Resources & Development. 5 (1): 13-24. ISSN 2409-2797. DOI: https://doi.org/10.17102/cnr.2018.02

Dorji, U. (2018). Study on diversity of Fishes in  Nyera Amari Chhu. ESIA report submitted to National Environment Commission Secretariat.  Kuensel Corporation Limited: Thimphu. ISBN 978-99936-994-0-8. https://www.researchgate.net/project/Study-on-diversity-of-Fishes-in-Nyera-AmariChhu  

Dorji, U. (2018). Analysing Carbon Stock in different forest and land uses in the context of Climate Variability and change in Western Bhutan. https://bhutanhimalayanclimatestudies.com/publications/analyzing-carbon-stock-indifferent-forest-and-land-uses-in-the-context-of-climate-variability-and-change-in-western-bhutan/  

Dorji, U. (2017, 10-12 May ). Brief account on freshwater biodiversity of Bhutan. Paper presented at the Second Bhutan Water Seminar-Water-Securing Bhutan's Future: “Towards Ensuring Water Security for Bhutan’s Future”,  Kuensel Corporation Limited: Thimphu. ISBN 978-99936-994-0-8. https://wwfasia.awsassets.panda.org/downloads/national_water_symposium_resolutions.pdf

Dorji, U. & Gurung, D. B. (2017). Aquatic Biodiversity of Bhutan. An Introduction to the Biodiversity of Bhutan: in the context of climate change and economic development. Kuensel Corporation Limited: Thimphu. ISBN 978-99936-994-0-8. https://www.researchgate.net/publication/352092371_Aquatic_Biodiversity_of_Bhutan  

Dorji, U. (2013). Invasive species in Ecological Habitats of National Parks in Bhutan, a review paper. https://www.scribd.com/document/174802072/INVASIVE-SPECIES-IN-ECOLOGICAL-HABITATS-OFNATIONAL-PARKS-IN-BHUTAN.  

Dorji, U. (2012). A pilot study on Carbon Footprint in Kanglung with particular reference to areas in and around Sherubtse College campus. Indo-Bhutan International Conference in Advances in Environmental Sciences (AES-2012).  Kuensel Corporation Limited: Thimphu. ISBN 978-99936-994-0-8. 

B. Editorial and Perspectives on Kuensel

1. Kuensel: 2025, January 11: Maximising the potential of groundwater resources in Bhutan.

2. Kuensel: 2024, October 25: The Silent Invasion: Microplastic Pollution in Bhutan's Urban Environment.

3. Kuensel: 2024, July 15: How safe is Roof-harvested Rainwater?

4. Kuensel: 2024, February 25: Microplastic Contamination of Bottled Water: Should we reconsider our sip?

5. Kuensel: 2024, January 13: The Growing Shadow of Emerging Contaminants in Drinking Water.

6. Kuensel: 2023, November 19: Considering Options to Construct Water Reservoir to address the drinking water shortage?

7. Kuensel: 2023, September 26: Reducing Risks from Flash Flood Events.

8. Kuensel: 2023, August 8: Conversion of urine into fertiliser.

9. Kuensel: 2023, July 8: Nutrient Enrichment in Alpine Wetlands - Case Study of Phobjikha.

10. Kuensel: 2023, March 18: How much water is stored in Thimphu’s drinking water catchments?

11. Kuensel: 2023, March 18: Drinking water quality concerns after Changjiji and Debsi fire.

12. Kuensel: 2023, February 27: Preventing disappearance of ‘holy water’ in Culturally Significant Sites.

13. Kuensel: 2023, January 31: Calling for water-sensitive urban designs for Bhutan’s Cities.

14. Kuensel: 2022, December 31: Promoting Sustainable Native Fish Population.

15. Kuensel: 2022, November 30: Water bugs are critical to our livelihood than we think.

16. Kuensel: 2022, October 30: New approaches to water source protection.

17. Kuensel: 2022, September 30: Proposing improvement in current septic tank systems for wastewater management.

18. Kuensel: 2022, August 31: Evaluating connectivity between precipitation & water systems for reviving drying water sources.

19. Kuensel: 2022, July 30: Managing toxic landfill leachates in dumpsites across Bhutan.

20. Kuensel: 2022, June 28: Understanding the linkage between snow-capped peaks to flow patterns of streams and rivers.

21. Kuensel: 2022, May 14: Challenges facing sustainable irrigation system in Bhutan.

22. Kuensel: 2022, April 4: Has Bhutan overlooked the transboundary GLOF threat so far?

23. Kuensel: 2022, March 29: Managing flash floods with localised river discharge modelling.

24. Kuensel: 2022, February 15: Abstraction of Groundwater in Mountainous Regions - Bhutan.

25. Kuensel: 2022, January 12: How safe is the treatment of our drinking water?