Two Decades of Proboscis Monkeys: A Conservation Success Story in Borneo
Kyoto University, Japan (2024–2025) | Master's Thesis Research
The flooded forests along Borneo's Kinabatangan River are home to one of the largest remaining populations of proboscis monkeys—a species found nowhere else on Earth. But how are they really doing? For my master's research, I set out to answer this question by looking back at more than 20 years of population data, creating one of the longest-term studies ever conducted on this charismatic species.
What We Did
This wasn't your typical fieldwork. Over several weeks, I conducted boat-based surveys along 223 kilometers of the Kinabatangan River scanning the riverbanks for proboscis monkeys. But the real work was just beginning. I combined my new observations with historical data stretching back to 2005 and 2014, creating a two-decade timeline of population changes alongside populations at three key monitoring sites. Using sophisticated statistical models—state-space model & GIS—I analyzed not just monkey numbers, but also how land use and forest cover had changed across their potential habitat from 2005 to 2025.
What We Discovered
The results surprised me—in the best way possible:
Good news for proboscis monkeys:
Overall population estimates in 2025 were higher than they were in 2005, suggesting the species is holding its own or even recovering in this region.
Stability matters:
Across the long-term monitoring sites, population sizes remained stable—no dramatic crashes, which is exactly what conservationists—including myself!—hope to see.
Forest is returning:
When I analyzed satellite imagery, I found that natural vegetation had actually increased over the study period. Importantly, there was little conversion of forest to oil palm plantations, particularly within protected areas (Wildlife Sanctuary and forest reserves).
Why It Matters
This research offers something rare in conservation: genuine hope. While proboscis monkeys remain endangered, our findings suggest that protection efforts in the Lower Kinabatangan are working. The stability and modest growth we documented likely reflect decades of conservation work—establishing protected areas, restoring degraded forest, and maintaining wildlife corridors.
But there's also a warning in these results. The fact that populations remained stable rather than booming tells us that the habitat can only support so many monkeys. Protecting every remaining patch of forest, and continuing to restore degraded areas, remains essential for their long-term survival.
For me, this project was a powerful reminder that conservation is a marathon, not a sprint. Two decades of monitoring—by multiple researchers across generations—allowed us to see the big picture. And that picture, while not perfect, is genuinely encouraging.
Disclaimer: This research represents my master's thesis work and is being prepared for peer-reviewed publication. The findings described here are based on analyses completed in 2025.
Parasites and Proboscis Monkeys: Health in a Changing World
Kyoto University, Japan (2024–2025) | Collaborative Research Project
Deep in the riverine forests of Sabah, Malaysian Borneo, lives one of the most unusual primates on Earth: the proboscis monkey. With their impressive noses, potbellies, and striking reddish-brown fur, these endangered monkeys face increasing pressure from habitat loss. In this study, I investigated an invisible but important aspect of their health—the parasites living inside their potbellies!
What We Did
Along the Menanggul River, a tributary of Sabah's Kinabatangan River, fecal samples from free-ranging proboscis monkey groups were collected. Back in the lab, we (led by Prof. Ikki) teamed up with parasite expert Dr. Liesbeth Frias to examine these samples under microscopes and identify any parasite eggs or larvae present. We then used Bayesian statistical models to explore whether factors like group size and habitat disturbance influenced infection rates by the parasites of the monkeys.
What We Discovered
The results were striking:
Widespread infection:
A remarkable 81.25% of the monkeys sampled showed evidence of parasite infection—meaning the vast majority carried at least one type of intestinal parasite.
Three main culprits:
We identified three dominant parasite groups, Trichuris sp., Strongyloides fuelleborni, and Oesophagostomum aculeatum and such parasites can also be found most wild primate populations—even humans!
Patterns in infection:
The number of parasites an individual carried wasn't random—it varied systematically with both how large their social group was and the condition of their surrounding habitat with variation depending on the species group of the parasites.
Why It Matters
Parasites are a normal part of ecosystem health, but changing conditions can upset the delicate balance between host and parasite. Proboscis monkeys are already endangered, and understanding how social factors and habitat disturbance affect their health gives conservation managers valuable insights. If larger groups or degraded habitats lead to higher parasite loads, these factors may need to be considered when planning protected areas or habitat restoration efforts.
This project was a fascinating glimpse into the hidden world of wildlife health—and a reminder that conservation isn't just about protecting habitat, but understanding the complex interactions that determine the animals' survivability.
📄 Read the preprint: doi.org/10.1101/2025.05.21.655443 This manuscript is currently under review for publication in a peer-reviewed journal.
Activity Patterns of Dusky Langurs in Penang Island, Malaysia
Universiti Sains Malaysia (2019–2020) | Bachelor's Thesis Research
In this study, I spent a year observing a fascinating group of dusky langurs—colobine, leaf-eating monkeys with dark silver fur and expressive spectacled-like faces and bright yellow babies—living along the coast of Teluk Bahang in Penang. What made this group special was their unique home: a patchwork landscape combining natural forest, a modified eco-tourism park, and beachfront areas.
What We Did
Every week, I followed the same group of langurs, carefully recording their behaviors and tracking their movements using GPS. My goal was to understand how these primates spend their time and whether younger monkeys behave differently from adults. Over the course of the study, I collected nearly 3,300 behavioral observations—1,716 from adults and 1,521 from juveniles.
What We Discovered
The data revealed some clear patterns:
Age matters:
Juveniles spent significantly more time moving and playing than adults—not surprising, given that young animals everywhere use play to develop strength and social skills.
Family matters:
Adults were more involved in carrying and caring for infants, a responsibility that juveniles haven't yet taken on.
Everything else:
When it came to feeding, resting, and social grooming, adults and juveniles behaved quite similarly.
Why It Matters
Understanding how wild primates use human-modified landscapes is increasingly important as development expands into natural habitats. This research shows that dusky langurs can adapt to areas with regular human activity, but also highlights which habitats they consider essential—information that can guide conservation efforts in Penang and beyond.
Disclaimer: This research represents my Bachelor thesis work with collaboration and field supervision by Langur Project Penang. All rights reserved.