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Email operates across computer networks, primarily the Internet, and also local area networks. Today's email systems are based on a store-and-forward model. Email servers accept, forward, deliver, and store messages. Neither the users nor their computers are required to be online simultaneously; they need to connect, typically to a mail server or a webmail interface to send or receive messages or download it.

Originally an ASCII text-only communications medium, Internet email was extended by Multipurpose Internet Mail Extensions (MIME) to carry text in other character sets and multimedia content attachments. International email, with internationalized email addresses using UTF-8, is standardized but not widely adopted.[1]

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The Simple Mail Transfer Protocol (SMTP) protocol was implemented on the ARPANET in 1983. LAN email systems emerged in the mid 1980s. For a time in the late 1980s and early 1990s, it seemed likely that either a proprietary commercial system or the X.400 email system, part of the Government Open Systems Interconnection Profile (GOSIP), would predominate. However, once the final restrictions on carrying commercial traffic over the Internet ended in 1995,[30][31] a combination of factors made the current Internet suite of SMTP, POP3 and IMAP email protocols the standard (see Protocol Wars).[32][33]

Many MTAs used to accept messages for any recipient on the Internet and do their best to deliver them. Such MTAs are called open mail relays. This was very important in the early days of the Internet when network connections were unreliable.[36][37] However, this mechanism proved to be exploitable by originators of unsolicited bulk email and as a consequence open mail relays have become rare,[38] and many MTAs do not accept messages from open mail relays.

Internet email messages consist of two sections, "header" and "body". These are known as "content".[41][42] The header is structured into fields such as From, To, CC, Subject, Date, and other information about the email. In the process of transporting email messages between systems, SMTP communicates delivery parameters and information using message header fields. The body contains the message, as unstructured text, sometimes containing a signature block at the end. The header is separated from the body by a blank line.

RFC 5322 specifies the syntax of the email header. Each email message has a header (the "header section" of the message, according to the specification), comprising a number of fields ("header fields"). Each field has a name ("field name" or "header field name"), followed by the separator character ":", and a value ("field body" or "header field body").

Email header fields can be multi-line, with each line recommended to be no more than 78 characters, although the limit is 998 characters.[43] Header fields defined by RFC 5322 contain only US-ASCII characters; for encoding characters in other sets, a syntax specified in RFC 2047 may be used.[44] In some examples, the IETF EAI working group defines some standards track extensions,[45][46] replacing previous experimental extensions so UTF-8 encoded Unicode characters may be used within the header. In particular, this allows email addresses to use non-ASCII characters. Such addresses are supported by Google and Microsoft products, and promoted by some government agents.[47]

RFC 3864 describes registration procedures for message header fields at the IANA; it provides for permanent and provisional field names, including also fields defined for MIME, netnews, and HTTP, and referencing relevant RFCs. Common header fields for email include:[50]

The To: field may be unrelated to the addresses to which the message is delivered. The delivery list is supplied separately to the transport protocol, SMTP, which may be extracted from the header content. The "To:" field is similar to the addressing at the top of a conventional letter delivered according to the address on the outer envelope. In the same way, the "From:" field may not be the sender. Some mail servers apply email authentication systems to messages relayed. Data pertaining to the server's activity is also part of the header, as defined below.

Internet email was designed for 7-bit ASCII.[59] Most email software is 8-bit clean, but must assume it will communicate with 7-bit servers and mail readers. The MIME standard introduced character set specifiers and two content transfer encodings to enable transmission of non-ASCII data: quoted printable for mostly 7-bit content with a few characters outside that range and base64 for arbitrary binary data. The 8BITMIME and BINARY extensions were introduced to allow transmission of mail without the need for these encodings, but many mail transport agents may not support them. In some countries, e-mail software violates RFC 5322 by sending raw[nb 2] non-ASCII text and several encoding schemes co-exist; as a result, by default, the message in a non-Latin alphabet language appears in non-readable form (the only exception is a coincidence if the sender and receiver use the same encoding scheme). Therefore, for international character sets, Unicode is growing in popularity.[60]

Most modern graphic email clients allow the use of either plain text or HTML for the message body at the option of the user. HTML email messages often include an automatic-generated plain text copy for compatibility.

Advantages of HTML include the ability to include in-line links and images, set apart previous messages in block quotes, wrap naturally on any display, use emphasis such as underlines and italics, and change font styles. Disadvantages include the increased size of the email, privacy concerns about web bugs, abuse of HTML email as a vector for phishing attacks and the spread of malicious software.[61]Some e-mail clients interpret the body as HTML even in the absence of a Content-Type: html header field; this may cause various problems.

Some web-based mailing lists recommend all posts be made in plain text, with 72 or 80 characters per line for all the above reasons,[62][63] and because they have a significant number of readers using text-based email clients such as Mutt.Various informal conventions evolved for marking up plain text in email and usenet posts, which later led to the development of formal languages like setext (c. 1992) and many others, the most popular of them being markdown.

Some Microsoft email clients may allow rich formatting using their proprietary Rich Text Format (RTF), but this should be avoided unless the recipient is guaranteed to have a compatible email client.[64]

Users can retrieve their messages from servers using standard protocols such as POP or IMAP, or, as is more likely in a large corporate environment, with a proprietary protocol specific to Novell Groupwise, Lotus Notes or Microsoft Exchange Servers. Programs used by users for retrieving, reading, and managing email are called mail user agents (MUAs).

When opening an email, it is marked as "read", which typically visibly distinguishes it from "unread" messages on clients' user interfaces. Email clients may allow hiding read emails from the inbox so the user can focus on the unread.[66]

Mail can be stored on the client, on the server side, or in both places. Standard formats for mailboxes include Maildir and mbox. Several prominent email clients use their own proprietary format and require conversion software to transfer email between them. Server-side storage is often in a proprietary format but since access is through a standard protocol such as IMAP, moving email from one server to another can be done with any MUA supporting the protocol.

Many current email users do not run MTA, MDA or MUA programs themselves, but use a web-based email platform, such as Gmail or Yahoo! Mail, that performs the same tasks.[67] Such webmail interfaces allow users to access their mail with any standard web browser, from any computer, rather than relying on a local email client.

Upon reception of email messages, email client applications save messages in operating system files in the file system. Some clients save individual messages as separate files, while others use various database formats, often proprietary, for collective storage. A historical standard of storage is the mbox format. The specific format used is often indicated by special filename extensions:

The URI scheme, as registered with the IANA, defines the mailto: scheme for SMTP email addresses. Though its use is not strictly defined, URLs of this form are intended to be used to open the new message window of the user's mail client when the URL is activated, with the address as defined by the URL in the To: field.[68][69] Many clients also support query string parameters for the other email fields, such as its subject line or carbon copy recipients.[70]

Many email providers have a web-based email client. This allows users to log into the email account by using any compatible web browser to send and receive their email. Mail is typically not downloaded to the web client, so it cannot be read without a current Internet connection.

The Internet Message Access Protocol (IMAP) provides features to manage a mailbox from multiple devices. Small portable devices like smartphones are increasingly used to check email while traveling and to make brief replies, larger devices with better keyboard access being used to reply at greater length. IMAP shows the headers of messages, the sender and the subject and the device needs to request to download specific messages. Usually, the mail is left in folders in the mail server.

Email has been widely accepted by businesses, governments and non-governmental organizations in the developed world, and it is one of the key parts of an 'e-revolution' in workplace communication (with the other key plank being widespread adoption of highspeed Internet). A sponsored 2010 study on workplace communication found 83% of U.S. knowledge workers felt email was critical to their success and productivity at work.[74] ff782bc1db