Publications

Pal, T. S., Nandi, S., Sarkar, R., & Bhadra, A., 2025. When Life Gives You Lemons, Squeeze Your Way Through: Understanding Citrus Avoidance Behaviour by Free-Ranging Dogs in India. Applied Animal Behaviour Science.

Roy, A., Lahiri, A., Nandi, S., Manchalwar, A., Siddharth, S., Abishek, J.V.R., Bulhan, I., Sengupta, S., Kumar, S., Chakravarty, T. and Bhadra, A., 2025. Ready, set, yellow! color preference of Indian free-ranging dogs. Animal Cognition.

Biswas,S., Ghosh,K.,Ghosh,S., Biswas A and Bhadra, A.,2024. What is in a scent? Understanding the role of scent marking in social dynamics and territoriality of free-ranging dogs. Behavioral Ecology and Sociobiology.

Biswas,S., Ghosh,K.,Sarkar,K., Benny, L., Katti, M. and Bhadra, A.,2024. A population-level study reveals hidden patterns in resting site choice of free-ranging dogs . Biological Journal of the Linnean Society.

Nandi,S., Mousumi Chakraborty, M., Lahiri,A., Gope,H.,Bhaduri, K.S. and Bhadra, A,2024. Free-ranging dogs quickly learn to recognize a rewarding person. Applied Animal Behaviour Science.

Biswas,S., Bhowmik,T., Ghosh,K., Roy, A., Lahiri, A., Sarkar,S., and Bhadra, A.,2024. Scavengers in the human-dominated landscape: an experimental study. Philosophical Transactions of the Royal Society B.

Bhattacharjee,D., Sau,S., Das,J and Bhadra,A,2024. Does novelty influence the foraging decisions of a scavenger, PeerJ.

Mondal,R., Protopopova, A and Bhadra, A.,2023. The human-animal bond and at-home behaviours of adopted Indian free-ranging dogs.  Applied Animal Behaviour Science. 

Sarkar, R., Bhowmick, A., Dasgupta, D., Banerjee, R., Chakraborty, P., Nayek, A., Sreelakshmi,R., Roy, A., Sonowal, R., Mandal, A.B. and Bhadra, A., 2023. Eating Smart: Free-ranging dogs follow an optimal foraging strategy while scavenging in groups. Frontiers in Ecology and Evolution.

Bhattacharjee, D. and Bhadra,A.,2021.  Adjustment in the point-following behaviour of free-ranging dogs - roles of social petting and informative-deceptive nature of cues. Animal Cognition.

Banerjee, A. and Bhadra, A., 2021. Time-activity budget of urban-adapted free-ranging dogs. acta ethologica, pp.1-10. 

Banerjee, A., Das, N., Dey, R., Majumder, S., Shit, P., Banerjee, A., Ghosh, N. and Bhadra, A., 2021. Power-laws in dog behavior may pave the way to predictive models: A pattern analysis study . Heliyon, p.e07243 .

Bhattacharjee, D. and Bhadra, A., 2021. Response to short-lived human overcrowding by free-ranging dogs . Behavioural Ecology and Sociobiology,75(7), pp.1-8

Bhattacharjee, D. and Bhadra, A., 2020. Humans Dominate the Social Interaction Networks of Urban Free-Ranging Dogs in India. Frontiers in psychology, 11, p.2153. 

Bhattacharjee, D., Sarkar, R., Sau, S., & Bhadra, A. 2020. Sociability of Indian free-ranging dogs (Canis lupus familiaris) varies with human movement in urban areas. Journal of Comparative Psychology. 

Bhattacharjee, D., Sau, S. and Bhadra, A., 2020. ‘Bolder’ together—response to human social cues in groups of free-ranging dogs. Behaviour, 157(3-4), pp.363-384. 

Bhattacharjee, D., Mandal, S., Shit, P., Varghese, M.G., Vishnoi, A. and Bhadra, A., 2020. Free-Ranging dogs are capable of utilizing complex human pointing cues. Frontiers in Psychology, 10, p.2818. 

Brubaker, L., Bhattacharjee, D., Ghaste, P., Babu, D., Shit, P., Bhadra, A. and Udell, M.A., 2019. The effects of human attentional state on canine gazing behaviour: a comparison of free-ranging, shelter, and pet dogs. Animal cognition, 22(6), pp.1129-1139. 

Banerjee, A. and Bhadra, A., 2019. The more the merrier: dogs can assess quantities in food-choice tasks. Current Science, 113891(117), p.6. 

Sarkar, R., Sau, S. and Bhadra, A., 2019. Scavengers can be choosers: A study on food preference in free-ranging dogs. Applied Animal Behaviour Science, 216, pp.38-44. 

Bhattacharjee, D., Sau, S. and Bhadra, A., 2018. Free-ranging dogs understand human intentions and adjust their behavioral responses accordingly. Frontiers in Ecology and Evolution, 6, p.232. 

Paul, M. and Bhadra, A., 2018. The great Indian joint families of free-ranging dogs. PloS one, 13(5). 

Bhattacharjee, D., Sau, S., Das, J. and Bhadra, A., 2017. Free-ranging dogs prefer petting over food in repeated interactions with unfamiliar humans. Journal of Experimental Biology, 220(24), pp.4654-4660. 

Bhattacharjee, D., Dasgupta, S., Biswas, A., Deheria, J., Gupta, S., Dev, N.N., Udell, M. and Bhadra, A., 2017. Practice makes perfect: familiarity of task determines success in solvable tasks for free-ranging dogs (Canis lupus familiaris). Animal cognition, 20(4), pp.771-776. 

Brubaker, L., Dasgupta, S., Bhattacharjee, D., Bhadra, A. and Udell, M.A., 2017. Differences in problem-solving between canid populations: Do domestication and lifetime experience affect persistence?. Animal Cognition, 20(4), pp.717-723. 

Bhattacharjee, D., Dev, N., Gupta, S., Sau, S., Sarkar, R., Biswas, A., Banerjee, A., Babu, D., Mehta, D. and Bhadra, A., 2017. Free-ranging dogs show age related plasticity in their ability to follow human pointing. PloS one, 12(7). 

Paul, M., Sau, S., Nandi, A.K. and Bhadra, A., 2017. Clever mothers balance time and effort in parental care: a study on free-ranging dogs. Royal Society open science, 4(1), p.160583. 

Paul, M. and Bhadra, A., 2017. Selfish pups: weaning conflict and milk theft in free-ranging dogs. PloS one, 12(2). 

Sen Majumder, S., Paul, M., Sau, S. and Bhadra, A., 2016. Denning habits of free-ranging dogs reveal preference for human proximity. Scientific reports, 6, p.32014. 

Paul, M., Sen Majumder, S., Sau, S., Nandi, A.K. and Bhadra, A., 2016. High early life mortality in free-ranging dogs is largely influenced by humans. Scientific reports, 6, p.19641. 

Bhadra, A., Bhattacharjee, D., Paul, M., Singh, A., Gade, P.R., Shrestha, P. and Bhadra, A., 2016. The meat of the matter: a rule of thumb for scavenging dogs?. Ethology Ecology & Evolution, 28(4), pp.427-440. 

Paul, M., Sen Majumder, S., Nandi, A.K. and Bhadra, A., 2015. Selfish mothers indeed! Resource-dependent conflict over extended parental care in free-ranging dogs. Royal Society open science, 2(12), p.150580. 

Sen Majumder, S. and Bhadra, A., 2015. When love is in the air: understanding why dogs tend to mate when it rains. PloS one, 10(12). 

Paul, M., Sen Majumder, S. and Bhadra, A., 2014. Grandmotherly care: a case study in Indian free-ranging dogs. Journal of ethology, 32(2), pp.75-82. 

Sen Majumder, S., Chatterjee, A. and Bhadra, A., 2014. A dog's day with humans–time activity budget of free-ranging dogs in India. Current Science, pp.874-878. 

Paul, M., Sen Majumder, S. and Bhadra, A., 2014. Selfish mothers? An empirical test of parent-offspring conflict over extended parental care. Behavioural processes, 103, pp.17-22. 

Sen Majumder, S., Bhadra, A., Ghosh, A., Mitra, S., Bhattacharjee, D., Chatterjee, J., Nandi, A.K. and Bhadra, A., 2014. To be or not to be social: foraging associations of free-ranging dogs in an urban ecosystem. Acta Ethologica, 17(1), pp.1-8. 

Bhadra, A. and Bhadra, A., 2014. Preference for meat is not innate in dogs. Journal of ethology, 32(1), pp.15-22.