Dinosaur T-rex EXCLUSIVE

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In 1892, Edward Drinker Cope found two vertebral fragments of a large dinosaur. Cope believed the fragments belonged to an "agathaumid" (ceratopsid) dinosaur, and named them Manospondylus gigas, meaning "giant porous vertebra", in reference to the numerous openings for blood vessels he found in the bone.[2] The M. gigas remains were, in 1907, identified by Hatcher as those of a theropod rather than a ceratopsid.[3]

Dinosaur T-rex

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Sue Hendrickson, an amateur paleontologist, discovered the most complete (approximately 85%) and largest Tyrannosaurus skeleton in the Hell Creek Formation on August 12, 1990. The specimen Sue, named after the discoverer, was the object of a legal battle over its ownership. In 1997, the litigation was settled in favor of Maurice Williams, the original land owner. The fossil collection was purchased by the Field Museum of Natural History at auction for $7.6 million, making it the most expensive dinosaur skeleton until the sale of Stan for $31.8 million in 2020.[11] From 1998 to 1999, Field Museum of Natural History staff spent over 25,000 hours taking the rock off the bones.[12] The bones were then shipped to New Jersey where the mount was constructed, then shipped back to Chicago for the final assembly. The mounted skeleton opened to the public on May 17, 2000, in the Field Museum of Natural History. A study of this specimen's fossilized bones showed that Sue reached full size at age 19 and died at the age of 28, the longest estimated life of any tyrannosaur known.[13]

Two isolated fossilized footprints have been tentatively assigned to T. rex. The first was discovered at Philmont Scout Ranch, New Mexico, in 1983 by American geologist Charles Pillmore. Originally thought to belong to a hadrosaurid, examination of the footprint revealed a large 'heel' unknown in ornithopod dinosaur tracks, and traces of what may have been a hallux, the dewclaw-like fourth digit of the tyrannosaur foot. The footprint was published as the ichnogenus Tyrannosauripus pillmorei in 1994, by Martin Lockley and Adrian Hunt. Lockley and Hunt suggested that it was very likely the track was made by a T. rex, which would make it the first known footprint from this species. The track was made in what was once a vegetated wetland mudflat. It measures 83 centimeters (33 in) long by 71 centimeters (28 in) wide.[22]

The teeth of T. rex displayed marked heterodonty (differences in shape).[50][51] The premaxillary teeth, four per side at the front of the upper jaw, were closely packed, D-shaped in cross-section, had reinforcing ridges on the rear surface, were incisiform (their tips were chisel-like blades) and curved backwards. The D-shaped cross-section, reinforcing ridges and backwards curve reduced the risk that the teeth would snap when Tyrannosaurus bit and pulled. The remaining teeth were robust, like "lethal bananas" rather than daggers, more widely spaced and also had reinforcing ridges.[52] Those in the upper jaw, twelve per side in mature individuals,[50] were larger than their counterparts of the lower jaw, except at the rear. The largest found so far is estimated to have been 30.5 centimeters (12 in) long including the root when the animal was alive, making it the largest tooth of any carnivorous dinosaur yet found.[53] The lower jaw was robust. Its front dentary bone bore thirteen teeth. Behind the tooth row, the lower jaw became notably taller.[50] The upper and lower jaws of Tyrannosaurus, like those of many dinosaurs, possessed numerous foramina, or small holes in the bone. Various functions have been proposed for these foramina, such as a crocodile-like sensory system[54] or evidence of extra-oral structures such as scales or potentially lips,[55][56][57] with subsequent research on theropod tooth wear patterns supporting such a proposition.[58]

Paul rejected the objections raised by critics, insisting that they are unwilling to consider that Tyrannosaurus might represent more than one species.[74] In a subsequent paper awaiting publication, Paul maintained the conclusion that Tyrannosaurus consists of three species. He pointed out that the criticism of the study naming T. imperator and T. regina only focused on two of the features used to distinguish the two new species (the number of small incisiform teeth and femur robustness), while the original study also compared the robustness of other bones as well (the maxilla, dentary, humerus, ilium and metatarsals). Furthermore, Paul argued that Tyrannosaurus can be separated into three different species based on the shape of knob-like bumps ('postorbital bosses') behind the eyes. Paul also argued that past research concluding that Tyrannosaurus only consists of one species (T. rex) has simply assumed that all Tyrannosaurus skeletons are a single species, and that many new dinosaur species have been named on the basis of fewer differences than he and his colleagues used when proposing T. imperator and T. regina.[75]

A study by Hutchinson and colleagues in 2011 corroborated the previous estimation methods in general, but their estimation of peak growth rates is significantly higher; it found that the "maximum growth rates for T. rex during the exponential stage are 1790 kg/year".[29] Although these results were much higher than previous estimations, the authors noted that these results significantly lowered the great difference between its actual growth rate and the one which would be expected of an animal of its size.[29] The sudden change in growth rate at the end of the growth spurt may indicate physical maturity, a hypothesis which is supported by the discovery of medullary tissue in the femur of a 16 to 20-year-old T. rex from Montana (MOR 1125, also known as B-rex). Medullary tissue is found only in female birds during ovulation, indicating that B-rex was of reproductive age.[96] Further study indicates an age of 18 for this specimen.[97] In 2016, it was finally confirmed by Mary Higby Schweitzer and Lindsay Zanno and colleagues that the soft tissue within the femur of MOR 1125 was medullary tissue. This also confirmed the identity of the specimen as a female. The discovery of medullary bone tissue within Tyrannosaurus may prove valuable in determining the sex of other dinosaur species in future examinations, as the chemical makeup of medullary tissue is unmistakable.[98] Other tyrannosaurids exhibit extremely similar growth curves, although with lower growth rates corresponding to their lower adult sizes.[99]

Over half of the known T. rex specimens appear to have died within six years of reaching sexual maturity, a pattern which is also seen in other tyrannosaurs and in some large, long-lived birds and mammals today. These species are characterized by high infant mortality rates, followed by relatively low mortality among juveniles. Mortality increases again following sexual maturity, partly due to the stresses of reproduction. One study suggests that the rarity of juvenile T. rex fossils is due in part to low juvenile mortality rates; the animals were not dying in large numbers at these ages, and thus were not often fossilized. This rarity may also be due to the incompleteness of the fossil record or to the bias of fossil collectors towards larger, more spectacular specimens.[99] In a 2013 lecture, Thomas Holtz Jr. suggested that dinosaurs "lived fast and died young" because they reproduced quickly whereas mammals have long life spans because they take longer to reproduce.[102] Gregory S. Paul also writes that Tyrannosaurus reproduced quickly and died young, but attributes their short life spans to the dangerous lives they lived.[103]

The discovery of feathered dinosaurs led to debate regarding whether, and to what extent, Tyrannosaurus might have been feathered.[104][105] Filamentous structures, which are commonly recognized as the precursors of feathers, have been reported in the small-bodied, basal tyrannosauroid Dilong paradoxus from the Early Cretaceous Yixian Formation of China in 2004.[106] Because integumentary impressions of larger tyrannosauroids known at that time showed evidence of scales, the researchers who studied Dilong speculated that insulating feathers might have been lost by larger species due to their smaller surface-to-volume ratio.[106] The subsequent discovery of the giant species Yutyrannus huali, also from the Yixian, showed that even some large tyrannosauroids had feathers covering much of their bodies, casting doubt on the hypothesis that they were a size-related feature.[107] A 2017 study reviewed known skin impressions of tyrannosaurids, including those of a Tyrannosaurus specimen nicknamed "Wyrex" (BHI 6230) which preserves patches of mosaic scales on the tail, hip, and neck.[5] The study concluded that feather covering of large tyrannosaurids such as Tyrannosaurus was, if present, limited to the upper side of the trunk.[104]

Only a single Tyrannosaurus specimen has been conclusively shown to belong to a specific sex. Examination of B-rex demonstrated the preservation of soft tissue within several bones. Some of this tissue has been identified as a medullary tissue, a specialized tissue grown only in modern birds as a source of calcium for the production of eggshell during ovulation. As only female birds lay eggs, medullary tissue is only found naturally in females, although males are capable of producing it when injected with female reproductive hormones like estrogen. This strongly suggests that B-rex was female and that she died during ovulation.[96] Recent research has shown that medullary tissue is never found in crocodiles, which are thought to be the closest living relatives of dinosaurs. The shared presence of medullary tissue in birds and other theropod dinosaurs is further evidence of the close evolutionary relationship between the two.[114] ff782bc1db