Peer Reviewed Articles

Fraga, J.K.R., S.S. Mendoza, A.B. Velasco, J.L. Estrada, D.E.M. Husana, F. S. Magbanua (2026). Diving deep: Trends and gaps in Philippine groundwater research (1982–2024). Journal of Hydrology: Regional Studies, 64. See here.

Reyes, N., J.V. Pleto, and D.E.M. Husana (2025). Ecological Implications of Isolated Microplastics from Mud Crab, Scylla serrata (Forsskål, 1775), and Its Estuarine Habitat in Pagbilao, Quezon Province, the Philippines. Philippine Journal of Science, 154, (6B): 1803-1819. See here.

Velasco A.B., F.S. Magbanua, J.S. del Rosario, J.L. Estrada, J.K.R. Fraga, J.P. Mondejar, J.F.J.F. Sapitan, I.P. Geroza, G.L. Tenoc, D.E.M. Husana (2024). Groundwater Quality Variations during Wet and Dry Seasons in Agricultural and Forested Areas in the Philippines. Water Resources, 51 (2): 293–S305. See here.

Tan, Heok Hui and D.E.M. Husana (2021). Barbodes pyrpholeos new species, the first cave-dwelling cyprinid fish in the Philippines, with redescription of B. montanoi (Teleostei: Cyprinidae). Raffles Bulletin of Zoology, 69: 309–323. See here. 

Husana, D.E.M (2020). Sundathelphusa prosperidad, sp. n. (Decapoda: Brachyura: Gecarcinucidae), a new species of cave-obligate freshwater crab from Mindanao Island, the Philippines, with notes on the conservation status of Philippine cave species. Journal of Cave and Karst Studies, 82 (3): 210–218 . See here. 

Husana, D.E.M., and P.K.L. Ng (2019). On the identity of Sundathelphusa philippina (von Martens, 1868) (Decapoda: Brachyura: Gecarcinucidae) from the Philippines, with descriptions of two new species. Zootaxa, 4585 (2): 315–331. See here.

Larson, H.K. and D.E.M. Husana (2018). A new species of the blind goby Caecogobius (Govioidei, Gobiidae, Gobinellinae) from a cave system in Mindanao, Philippines. Ichthyological Research. See here.

Ishikawa, M., N.O. Ogawa, N. Ohkouchi, D.E.M. Husana and T. Kase (2017). Stable carbon isotope compositions of foot tissue, conchiolin opercula and organic matrix within shells in two marine snails, Naticidae Polinices mammilla and Ampullinidae Cernina fluctuata, from a seagrass meadow in the Philippines. Geochemical Journal, 51(3): 241–250. See here.

Husana, D.E.M., J.C.E. Mendoza and T. Kase (2015). Two new species of freshwater crab genus Sundathelphusa (Decapoda: Brachyura: Gecarcinucidae) from Negros Island, Philippines. Raffles Museum of Biodiversity Research, 63: 226–236. See here.

Husana, D.E.M., M. Yamamuro and P.K.L. Ng (2014). Two new species of freshwater crabs of the genus Sundathelphusa Bott, 1969 (Decapoda: Brachyura: Gecarcinucidae) from caves in Luzon, Philippines. Zootaxa, 3815 (4): 565–574. See here.

Husana, D.E.M. and T. Kikuchi (2013). Concealed environmental threat in the coastal region requires persistent attention: The Panglao Island, Philippines example. Journal of Environmental Protection, 4:1149–1156. See here.

Husana, D.E.M., T. Kase and P.K.L. Ng (2013). On a new species of freshwater crab, Ovitamon Ng & Takeda 1992 (Crustacea, Brachyura, Potamidae) from Panay Island, Philippines. Raffles Museum of Biodiversity Research, 61(2):651–655. See here.

Husana, D.E.M. and M. Yamamuro (2013). Groundwater quality on karst regions in the Philippines. Limnology, 14(3): 293–299. See here. 

Husana, D.E.M., J.C.E. Mendoza and T. Kase (2013). New species of false spider crab, Elamena H. Milne Edwards, 1837 (Crustacea: Decapoda: Brachyura: Hymenosomatidae), from Davao Gulf, Philippines. Zootaxa, 3616(5):495–600. See here. 

Husana, D.E.M., S.H. Tan and T. Kase (2011). A new genus and species of anchialine Hymeosomatidae (Crustacea, Decapoda, Brachyura) from Samar, Philippines. Zootaxa, 3109:49–59. See here 

Husana, D.E.M., T. Naruse, and T. Kase (2010). A new species of the genus Karstarma (Crustacea: Decapoda: Brachyura: Sesarmidae) from anchialine caves in the Philippines. Raffles Museum of Biodiversity Research 58(1):51–55. See here.

Husana, D.E.M., T. Naruse, and T. Kase (2009). Two new cavernicolous species of the genus Sundathelphusa from Western Samar, Philippines (Decapoda: Brachyura: Parathelphusidae). Journal of Crustacean Biology 29(3): 419–427. See here.

Cai, Y. and D.E.M. Husana (2009). Cave shrimps of the genus Edoneus Holthuis, 1978, from Luzon, the Philippines, with descriptions of three new species (Drustacea: Decapoda: Atyidae). Raffles Museum of Biodiversity Research 57(1):51–63. See here.

Book, Lab Manual, etc.

Fraga, J.K.R.,… D.E.M. Husana, et al. Bukas Booklet: A Guide to Philippine Stygofauna. Institute of Biology, University of the Philippines Diliman, 65 pages. 2024.

Fraga, J.K.R.,… D.E.M. Husana, et al. TAPAYAN: Sinalok na Kaalaman Tungkol sa Groundwater. Institute of Biology, University of the Philippines Diliman, 36 pages. 2024.

Banaay, C.G.B., D.E.M. Husana and F.S. Maranan. Science Literacy for All: Earth Science. Textbook for senior high school. ABIVA Publishing House, Inc. Philippines, 331 pages. 2019.

Husana, D.E.M. Contribution Box 2.5 p. 76. In: Philippine Caves: Beneath the Earth’s Surface – Conservation and Management. Quezon City: Department of Environment and Natural Resources-Biodiversity Management Bureau. 294 pages. 2017.

Fajardo, A.M., R.T. Beringuela, T.L.A. Honrade and D.E. Husana. Exploring Pollution in Biological Systems. Laboratory Manual. University of the Philippines Los Baños, 60 pages. 2015.

Conference Presentations

Reyes, N., J.V. Pleto, and D.E.M. Husana. Microplastics from Scylla serrata (Forsskal, 1775), and its estuarine habitat in Pagbilao Mangrove Forest. Poster. BIOTA, UPLB, Philippines, 24 January 2026.

DeLuna C.A.P. and D.E.M. Husana. Morphotaxonomic review of the freshwater crab Sundathelphusa Bott, 1969, inhabiting Mt. Makiling, Luzon, the Philippines. International Symposium & 42nd Annual Meeting of the Association of Systematic Biologists of the Philippines, UST, Manila, Philippines, 26–28 May 2025.

Martinez, A.,…D.E. Husana,…et al. Stygofauna Mundi: a comprehensive global biodiversity database of groundwater-related habitats across marine and freshwater realms. 26th International Conference on Subterranean Biology and 6th International Symposium on Anchialine Ecosystems. Cagliari (Sardinia), Italy, 9–14 September 2024.

Fraga, J.K.R., J.L. Estrada, D.E.M. Husana and F.S. Magbanua. Temporal variation of stygofauna richness from selected groundwater sources in the Philippines. Contributed poster paper (3rd place, poster award). 4th Philippine Symposium on Freshwater Biodiversity and Ecosystems in conjunction with 5th International Conference on Tropical Limnology 3rd General Assembly Meeting of the Philippine Society for Freshwater Science, Silliman University, Dumaguete City, Philippines, 13–15 November 2024.

Velasco, S.B., J.K.R. Fraga,…D.E.M. Husana, et al. The Philippine Groundwater Health Index: A tool in assessing groundwater health in agricultural areas. Contributed oral paper. 4th Philippine Symposium on Freshwater Biodiversity and Ecosystems in conjunction with 5th International Conference on Tropical Limnology 3rd General Assembly Meeting of the Philippine Society for Freshwater Science, Silliman University, Dumaguete City, Philippines, 13–15 November 2024.

Estrada, J.L., D.E. Husana and F.S. Magbanua. Taxonomic Abundance and Richness of Stygofauna Communities in the Groundwater within Agricultural Areas in Cebu, Philippines. Contributed oral paper. Freshwater Sciences 2023, Brisbane, Australia, 3–7 June 2023.

Macandog, M.M.,…D.E.M. Husana…C.G.B. Banaay. Blended, Relevant and Upskilled: Jumpstarting the Redesign of an Undergraduate Ecology Curriculum, 2022 AUN-EEC Conference (Oral, International), online 14–15 September 2022

Banaay, C.G.B., F.S. Maranan and D.E.M. Husana. Assessment of the Biomechanical filtration ability of Asiatic clams (Corbicula fluminea) from the south bay of Laguna Lake, Philippines. Contributed oral paper. 18th World Lake Conference, Guanajuato, Mexico, (online) 9–11 November 2021.

Maranan, F.S., C.G.B. Banaay and D.E.M. Husana. Leveraging project-based learning for ecological education: Evaluation of a Corbicula fluminea project as a medium for teaching ecological concepts. Oral presentation. 2021 ASEAN University Network-Ecological Education and Culture Conference 15 September 2021.

Macandog, DM, D.E.M. Husana, M.G.P. Tiquio, N.K. Torreta, E.R.C. DeChavez, A.R. Salvacion, J.G. Campang, J.V.R. Pleto, N.J.A Larida. Ecological Assessment of Laguna Lake Watershed, Water Quality and Biodiversity Resources. Oral presentation. 35th Congress of the International Society of Limnology (SIL 2021), Gwangju, Korea. 22-27 August 2021.

Banaay, C.G.B.B., A. Fajardo, F.S. Maranan, M.A.F. Rabena and D.E.M. Husana (2020). Refining the First Steps: Transcending Ecological Knowledge and Understanding into Lifelong Habits and Behaviors. Oral presentation. First AUN-EEC Conference on Ecological Education and Culture: Cultivating Ecological Mindsets and Lifestyles, online, 17 November 2020. 

Husana, D.E.M. and M. Yamamuro (2018). Trophic structure in a tropical cave ecosystem: surface-subsurface ecosystem interaction and implications for conservation. Oral presentation. European Conference of Tropical Ecology, Paris, Frarnce, 26-29 March 2018. Abstract S18-O04.

Quitoriano, J.P.S., M.K.O. Paler, P.S. Tiongson, Q.B. Amante and D.E.M. Husana (2017). Mortality and negative geotaxis behavior of Drosophila melanogaster Meigen, 1830 in response to low concentration of Cypermethrin. Oral paper. 52nd Biology Teachers Association of the Philippines (BIOTA). 6-8 April 2017, Philippine Normal University, Philippines.

Perez, C.A.G., F.S. Maranan, C.G.B. Banaay and D.E.M. Husana (2017). Initiatives, Opportunities, and Challenges in Creating a Management Plan for Ecotourism. Oral paper. International Conference of European and Asian Researchers and Educators on Lifelong Learning for Sustainable Development. 12-14 May 2017, Boracay Island, Philippines. 

Cunanan, D.J. and D.E.M. Husana (2016). Phylogeography of Philippine Sundathelphusa in surface-subsurface habitats: implications for conservation. Oral paper. Conservation Asia 2016: ACB-ATBC Asia Pacific Chapter. 29 June–3 July 2016, National University of Singapore, Singapore.

Husana, D.E.M., T. Haga and T. Kase (2016). Phylogeography of the crab genus Sundathelphusa: history of extensive migration, cave colonization and refugia in the Philippines. Oral Presentation. 23rd International Conference on Subterranean Biology. 12–17 June 2016, University of Arkansas, Fayetteville, Arkansas, USA.

Cunanan, D.J. and D.E.M. Husana (2016). Phylogeography, haplotype diversity and niche differentiation among freshwater crab Sundathelphusa (Decapoda: Brachyura: Gecarcinucidae) in the subterranean habitat of Quezon, Philippines. Oral Presentation. 23rd International Conference on Subterranean Biology. 12–17 June 2016, University of Arkansas, Fayetteville, Arkansas, USA.

Mendoza, R.M., D.E.M. Husana and C.G.B. Banaay (2014). Influence of nutrient load on the aquatic leaf litter decomposer community. Oral Presentation. 14th National Scientific Meeting of the Philippine Society for the Study of Nature. 20–24 May 2014, Benguet State University, Benguet, Philippines.

Husana, D.E.M., M. Yamamuro and T. Kase (2012). Colonization and speciation of cave animals in the Philippines. Poster Presentation. 2012 American Geophysical Union Fall Meeting. 3–7 Dec. 2012, Moscone Convention Center, Howard Street, Between Third & Fifth Sts., San Francisco, California, USA. Abstract B51A-0479.

Husana, D.E.M., T. Kase and M. Yamamuro. Caves in the Philippines: what should we know and why should we care? Oral presentation. 21st Annual Biodiversity Symposium of the Wildlife Conservation Society of the Philippines (WCSP).17–20 April 2012, Dasmariñas, Cavite, Philippines.

Husana, D.E.M., T. Kase and M. Yamamuro (2011). Cave ecology in the Philippines, a conservation perspective: linking surface and subsurface ecosystems. Oral presentation. 2011 International Congress for Conservation Biology. 5–9 December 2011, Auckland, New Zealand. O.243-139023.

Husana, D.E.M. and M. Yamamuro (2011). Unforeseen human impacts on groundwater in karst regions: observations and pollution detection in the Philippine caves. Oral presentation. International Conference on Environmental Pollution and Remediation (ICEPR’11), Ottawa, Ontario, Canada, 17–19 August 2011. Abstract 236.

Husana, D.E.M., T. Haga, T. Kase and M. Yamamuro (2010). Phylogeography, cave invasion and diversification of the Philippine Sundathelphusa (Decapoda: Brachyura: Parathelphusidae). Poster presentation. 2010 American Geophysical Union Fall Meeting. 13–17 Dec. 2010, Moscone Convention Center, Howard Street, Between Third & Fifth Sts., San Francisco, California, USA. B33F-045, p. 168.

Husana, D.E.M., T. Kase and M. Yamamuro (2009). Biodiversity of cavernicolous crabs and conservation of cave habitats in the Philippines. Oral presentation. The Crustacean Society summer meeting and 47th Carcinological Society of Japan annual meeting. 20–24 Sept. 2009, Tokyo University of Marine Science and Technology, Tokyo, Japan. G-A9, p.12.

Husana, D.E.M., T. Kase and M. Yamamuro (2009). Conservation of cave biodiversity, urbanization and sustainable use of groundwater resources. Poster presentation. Student Summit for Sustainability (S3) and Alliance for Global Sustainability (AGS) annual meeting. 26–29 Jan. 2009, ETH Zurich, Switzerland.

Husana, D.E.M., T. Kase and M. Yamamuro (2008). Biodiversity of cavernicolous crabs and conservation of cave habitats in the Philippines. Oral presentation. 19th International Symposium of Subterranean Biology. 21–26 Sept. 2008, Western Australian Museum Maritime precinct, Fremantle, Western Australia. p. 49.

Husana, D.E.M., T. Kase and T. Naruse (2007). An unusual troglobitic freshwater crab of the genus Sundathelphusa (Decapoda: Brachyura: Parathelphusidae) from caves in Samar, Philippines. Poster presentation. 45th annual meeting of Carcinological Society of Japan. 13–14 Oct. 2007, Faculty of Marine Science, Tokyo University of Marine Science and Technology, Tokyo, Japan. p. 56. 

Husana, D.E.M., T. Kase, and T. Naruse (2007). New cavernicolous freshwater crabs from Samar, Island, Philippines. Poster presentation. International Workshop on tropical island biodiversity: across land and sea, 25–26 Sept. 2007, National University of Singapore, Singapore. p.27.

Invited Talks and Lectures

Biodiversity and Ecology of the Philippine Cave Fauna. Philippine Speleological Society National Cave Congress. 9-13 April 2026. Mabinay, Negros Oriental, Philippines.

Appreciating Life from a Scientific Perspective: Re-exploring Freshwater Crabs and Mosses in the Cordillera. 27 October 2023. Benguet State University, Philippines.

Ecological Values of Philippine Terrestrial Caves, Philippine Cave and Karst Digital Forum, Session 3. 16 October 2020. Department of Environment and Natural Resources–Biodiversity Management Bureau, Philippines.

Biodiversity Assessment and Monitoring System (BAMS), Online Lecture Series—Cave Aquatic Fauna. 14 October 2020. Department of Environment and Natural Resources–Biodiversity Management Bureau, Philippines. 

The paradox of urbanization: understanding the concealed threat of groundwater deterioration. Sustainable Urban Development Program. 6-10 September 2018. Southeast Asian Regional Center for Graduate Study and Research in Agriculture (SEARCA), University of the Philippines Los Baños, College, Laguna, Philippines.

Trade-offs in the dark: unique adaptive features of cave aquatic fauna. 2nd Philippine Symposium on Freshwater Biodiversity and Ecosystems. 11­–14 December 2018. Institute of Biology, University of the Philippines Diliman, Quezon City, Philippines.

Cave Biology: Life in the perpetual darkness of the Philippine caves. 17th Philippine Speleological Society National Cave Congress. 24­–28 April 2017. Roxas, Zamboanga del Norte, Philippines.

Freshwater crab Sundathelphusa: model of successful dispersal and colonization? 1st Philippine Symposium on Freshwater Biodiversity and Ecosystems. 7­–10 June 2016. University of Santo Tomas, Manila, Philippines.

Biology and Ecology of Caves in the Philippines. College of Natural Sciences and Mathematics Seminar. 1 July 2014. MSU AVR Graduate School, Mindanao State University, General Santos City, Philippines.

What lies beneath: linking surface and subterranean ecosystems. 13th Graduate Research Forum, Science and Technology Advisory Council–Japan Chapter. 13 March 2010. TIEC, Odaiba, Japan.

Cave and Cave Life. 27 April 2009. Municipal Government of Prosperidad, Agusan del Sur, Philippines.

Cave Biodiversity and its Conservation. Lecture forum. 2 February 2004. Mindanao State University, General Santos City, Philippines.

Views: Ecology, belief, history and Arts in Holy Mountain - a colloquium with four views in Mt. Banahaw. 21–23 January 2004. Nawawalang Paraiso Resort, Tayabas, Quezon, Philippines.

Cave Biodiversity. Department of Environment and Natural Resources–Protected Areas and Wildlife Management Bureau National Training on Cave Management. 13–21 May 2003. Tuguegarao City, Philippines.

Dark Ecosystem. Natural Science Department, College of Sciences, seminar. 20 September 2001. Adamson University, Manila, Philippines.