Professor at National Taiwan University

Associate editor for Frontiers in Marine Science (Section: Marine Evolutionary Biology, Biogeography and Species Diversity)

Email:  wjchen.actinops[at]gmail[dot]com; wjchen1971[at]ntu[dot]edu[dot]tw

FB: https://www.facebook.com/MBPGLab/

Mailing address:
Room 301, Institute of Oceanography, National Taiwan University
No.1 Sec. 4 Roosevelt Rd., Taipei 10617, Taiwan

Phone: 886 (0)2-3366-1630; 886 (0)2-2363-6040 ext.301 Fax: 886 (0)2-2363-7062


Aug 2017 - Present

Full Professor at Institute of Oceanography and Department of Life Science (joint appointment), National Taiwan University, Taiwan.

Aug 2015 - July 2017

Associate Professor (joint appointment) at Department of Life Science, National Taiwan University, Taiwan

Dec 2017 - Feb 2018

Visiting Associate Professor at the Kyoto University Museum, Japan

Aug 2013 - July 2017

Associate Professor at Institute of Oceanography, National Taiwan University, Taiwan.

Aug 2009 - July 2013

Assistant Professor at Institute of Oceanography, National Taiwan University, Taiwan.

Feb 2006 - June 2009
Postdoctoral Research Fellow at Saint Louis University, USA (with Prof. Richard Mayden).

Oct 2003 - Dec 2005
Contractor-Researcher at INRA, Bordeaux, France (with Dr. Marie-France Corio-Costet).

Nov 2002 - May 2003
Postdoctoral Research Fellow at University of Konstanz, Germany (with Prof. Axel Meyer).

Feb 2001 - Oct 2002
Postdoctoral Research Fellow at University of Nebraska, USA (with Dr. Guillermo Ortí).

Feb 2002 & Feb 2006

Qualified as “maîtres de conférences” (assistant professor) in section 68 “Biologie des organismes” in France.



2001. Ph.D. Biological Sciences- Biodiversity: Genetics, History and Mechanisms of Evolution at University of Paris 6 (Université Pierre et Marie Curie), France.

1997. DEA (equivalent to master degree in USA system):Biodiversity: Genetics, History and Mechanisms of Evolution at University of Paris 6 (Université Pierre et Marie Curie), France.

1993. Bachelor of Science at Department of Zoology, Fishery Division, National Taiwan University, Taiwan.


My work and general research interests concern the application of molecular and computational tools within a phylogenetic framework to interpret organismal diversity (in fish particularly) and to understand mechanisms of evolution. Currently, I'm emphasizing on marine phylogenomics (see Chen and Mayden 2010Chen, Samadi, and Chen 2018), exploration of marine biodiversity (see Lee et al. 2019; Chen et al. 2016, 2017), biogeography (especially, in studying the origin of the Coral Triangle’s extreme concentration of marine biodiversity in relation to geological, climate and evolutionary events), and evolution of deep-sea metazoans (see Lee et al. 2019).

Media interview (海洋之窗-台灣海洋向前行): Youtube link 1, 2, 3, & 4


  • Biogeography [4, 7, 20, 26, 28, 31, 35, 42, 46, 72, in progress]
  • Biodiversity exploration from the Indo-West Pacific: shallow water or reef associated fauna [5, 9, 14, 17, 19-20, 24, 30, 36, in progress]; deep-sea fauna [2-3, 5, 10-12, 21-22, 29, 43, in progress]
  • Mitochondrial genomics of mollusks [5, 22, in progress] and ray-finned fishes [16, 33, 66]
  • High-level phylogeny of ray-finned fishes, e.g., Elopomorpha [26, 37], Acanthomorpha [33, 74]), and others [4, 16, 34-35, 37-39, 41, 69, 71]
  • Cypriniformes tree of Life (CToL) project [18, 32, 48-49, 51, 53, 58-60; 62-67]
  • Molecular systematics of the Teleostei with focus on 
- Pleuronectiformes [4, 33, 37, 41, in progress]
- Lethrinidae [24, 40, 45, 56-57, in progress]
Lutjanidae [9, in progress]
- Mugilidae [48-49, in progress]
- Nemipteridae [19, in progress]
- Sciaenidae [17, 28, in progress]

- Pomacentridae [20, 30-31, in progress]
Clupeoidei [13, 15-16]
- Gerreidae [69]
- Antarctic notothenioids [75-76]
  • Phylogenomics and evolutionary genomics [6, 8, 23, 25, 27, 54-55, 71]
  • Coral research [1, 25]
  • Development of nuclear gene markers for studying phylogenetics, systematics, and evolution of ray-finned fishes [60, 73-74] (see also this page)
  • Mechanisms of evolution of fungicide resistance in populations of grapevine downy mildew [52, 61, 68, 70]


    • Personal research funding from Ministry of Science and Technology, Taiwan (2019 – 2021) (MOST 108-2611-M-002 -012 -MY2) (4,022,000 NTD). Project title: Biodiversity exploration in the deep sea (second phase): Usage of emerging technologies and integrated approach to meet new challenges
    • Personal research funding from Ministry of Science and Technology, Taiwan (2018 – 2019) (MoST 07-2611-M-002-007-) (1,686,000 NTD). Project title: Biodiversity exploration in the deep sea


    • Personal research funding from Ministry of Science and Technology, Taiwan (2015 – 2018) (MoST 104-2611-M-002-002-MY3) (6,459,000 NTD). Project title:  ignificance of the Indo-West Pacific barrier(s) to marine species distribution and evolution: emperor breams (Perciformes: Lethrinidae) as a case study
    • Bilateral cooperation research funding from National Science Council, Taiwan  (2013 – 2016) (NSC 102-2923-B-002 -001 -MY3) (9,166,000 NTD) and ANR, France. Project title: Taiwan France marine diversity exploration and evolution of deep-sea fauna (TFDeepEvo)
    • Personal research funding from MOST, Taiwan (2012 – 2015) (NSC 101-2611-M-002 -016 -MY3) (5,966,000 NTD). Project title: Evolution and marine biogeography of fish of Taiwan and South China Sea islands in an Indo-West Pacific context
    • Subcontract of the cooperation research funding from National Science Foundation, USA (2013 – 2014) (30,800 USD). Project title: Collaborative PBI: All Cypriniformes Species: Phase II of an Inventory of the Otophysi
    • Personal research funding from MOST, Taiwan (2010-2012) (NSC 99-2611-M-002-001-MY2) (4,855,000 NTD). Project title: Marine phylogenomics: a case study for ray-finned fishes


    Sept. 2019
    New Caledonia in Central Pacific Ocean (Campaign: KANADEEP 2), with R/V Atalante, France
    July/Aug. 2019
    E of Taiwan/Luzon Strait Expedition 2019 (Campaign: Sakizaya 2019), with R/V OR1, Taiwan
    April 2019
    Scattered Islands (îles d'éparses) in W. Indian Ocean (Campaign: DHEEP), with R/V Marion Dufresne (link; PDF)
    June 2018
    NE of Taiwan Expedition 2018 (Campaign: Kavalan 2018), with R/V OR1, Taiwan
    July 2015
    ZhongSha/DongSha Exploration 2015 (Campaign: ZhongSha 2015), with R/V OR1, Taiwan
    May 2014
    DongSha Exploration 2014 (Campaign: DongSha 2014), with R/V OR1, Taiwan
    Jan. 2014
    Nan-Hai Marine Biodiversity Exploration 2014 (Campaign: NanHai 2014), with R/V OR5, Taiwan
    May 2013
    South and SE of Taiwan in West Pacific Ocean (Campaign: TAIWAN 2013), with R/V OR1 & 3, Taiwan
    Nov. 2012
    Biodiversity expedition in Madang, Papua New-Guinea (Program: Our Planet Reviewed)
    Sept. 2011
    New Caledonia in Central Pacific Ocean (Campaign: EXBODI), with R/V ALIS, IRD (see news)
    April 2011
    Scattered Islands (îles d'éparses) in W. Indian Ocean, with R/V Marion Dufresne (see media news)


    During my doctorate study (at Laboratoire d’Ichtyologie générale et appliquée and Service commun de systématique moléculaire, Muséum National d’Histoire Naturelle (MNHN) à Paris under the supervision of Dr. Guillaume Lecointre), my research was focused on a systematics project of acanthomorph fishes, the most diverse group of vertebrates. My work was the first attempt to summarize multi-locus data for an extensive taxonomic sampling of acanthomorphs [74]. I also worked on small-scale phylogeny of higher notothenioids (Antarctic acanthomorphs, composed of the families Harpagiferidae, Artedidraconidae, Bathydraconidae, and Channichthyidae) based on two mitochondrial genes [75-76]. The comparison with phylogeography and ecology of included channichthyid taxa allowed proposing putative speciation models and correlations with paleo-events of Antarctic Ocean [76].

    I embarked on my first postdoctoral training at the University of Nebraska-Lincoln to further horn my skills on molecular techniques and molecular systematics of fishes [73]. Later, I had a short stint as a postdoctoral fellow at University of Konstanz in Germany where I worked on projects of EVODEVO (evolutionary development) in cichlids and phylogenomic analysis of fish model systems [71]. Besides, a collaboration work (with Dr. Jing Luo) on molecular systematics and biogeography of Oriental Voles (Genus Eothenomys, Muridae, Mammalia) was accomplished [72].

    I also expand my extensive experience in Molecular Systematics studies to the investigations of micro-evolution and Molecular Ecology using 
    grapevine downy mildew (
    Plasmopara viticola) as model system [52,61,68,70]. This research activity was conducted at French National Institute for Agronomic research (INRA, Bordeaux).

    Prior to my current position, I have been working on the US NSF funded AToL study on the Cypriniformes Tree of Life (CToL) at St. Louis University [58-60; 62-67].



    1. CHEN, C.-J., CHEN, W.-J., SHIKINA, S., DENIS, V., & CHANG, C.-F. (2020) The plasticity of gonad development of sexual reproduction in a scleractinian coral, Porites lichen. General and Comparative Endocrinology 285:113270 (link).

    2. OGINO, A., LEE, S.-H., CHEN, W.-J., & MATSUNUMA, M. (2020) Chelidoperca cerasina sp. nov., a new perchlet (Perciformes: Serranidae) from the southwest Pacific Ocean. Ichthyological Research 67, 117-132 (link).

    3. LEE, S.-H., LEE, M.-Y., MATSUNUMA, M., & CHEN, W.-J. (2019) Exploring the phylogeny and species diversity of Chelidoperca (Teleostei: Serranidae) from the western Pacific Ocean by an integrated approach in systematics, with descriptions of three new species and a redescription of C. lecromi Fourmanoir, 1982. Frontiers in Marine Science 6:465 (link).

    4. CAMPBELL, M.A., CHANET, B., CHEN, J.-N. LEE, M.-Y., & CHEN, W.-J. (2019) Origins and relationships of the Pleuronectoidei:  Molecular and morphological analysis of living and fossil taxaZoologica Scripta 48(5), 640-656 (Link).

    5. FRICKE, R., ALLEN, G.R., AMON, D., ANDREFOUET, S., CHEN, W.-J., KINCH, J., MANA, R., RUSSELL, B.C., TULLY, D., & WHITE, W.T. (2019) Checklist of the marine and estuarine fishes of New Ireland Province, Papua New Guinea, western Pacific Ocean, with 810 new records. Zootaxa 4588(1), 1–360 (link).

    6. LEE, H., CHEN, W.-J.SAMADI, S., PUILLANDRE, N., TSAI, M-H, AZNAR-CORMAN,L. & SAMADI, S. (2019) Incorporation of deep-sea and small-sized species provides new insights into gastropods phylogeny. Molecular Phylogenetics and Evolution 135, 136–147 (link).

    7. LEE, H.⁠ , LIN⁠, J.-P.⁠ , LI⁠ , H.-C., CHANG, L.Y., LEE⁠, K.S., LEE, S.-J. , CHEN, W.-J.⁠, SANKAR, A., & KANG, S.-C. (2019) Young colonization history of a widespread sand dollar (Echinodermata; Clypeasteroida) in western Taiwan. Quaternary International 528(10), 120-129 (link).


    8. CHEN, J.-N., SAMADI, S., & CHEN, W.-J. (2018) Rhodopsin gene evolution in early teleost fishes. PLoS ONE, 13(11): e0206918 (link).

    9. BAKAR, A.A., ADAMSON, E.A.S., JULIANA, L.H., NOR MOHD S.A., CHEN, W.-J., MAN, A., et al. (2018) DNA barcoding of Malaysian commercial snapper reveals an unrecognized species of the yellow-lined Lutjanus (Pisces:Lutjanidae). PLoS ONE, 13(9): e0202945 (link). 

    10. OKAMOTO, M., CHEN, W.-J., & SHINOHARA, G. (2018) Epigonus okamotoi (Perciformes: Epigonidae), a junior synonym of E. draco, with new distributional records for E. atherinoides and E. lifouensis in the West Pacific. Zootaxa 4476 (1): 141–150 (link).

    11. WANG, S.-Y., CHEN, J.-N., RUSSELL, B.C., & CHEN, W.-J. (2018) First record of Gauguin’s blunt-nose lizardfish, Trachinocephalus gauguini Polanco, Acero & Betancur 2016 (Teleostei: Synodontidae) outside the Marquesas Archipelago. Zootaxa 4476 (1): 151–156 (link).

    12. TONGBOONKUA, T., LEE, M.-Y., & CHEN, W.-J., (2018) A new species of sinistral flatfish of the genus Chascanopsetta (Teleostei: Bothidae) from off Papua New Guinea, western Pacific Ocean. Zootaxa 4476 (1): 168–181 (link).

    13. LAVOUÉ, S., RATMUANGKHWANG, S., HO, H.-C., CHEN, W.-J., SITI AZIZAH, M.N. (2018) Swimbladder evolution of longfin herrings (Pristigasteridae, Teleostei). Zoological Studies 57:39 (link).


    14. CHEN, W.-J., MIKI, R., & BORSA, P. (2017) Gymnocranius obesus, a new large-eye seabream from the Coral Triangle. Comptes Rendus Biologies 340, 520-530 (link).

    15.  LAVOUÉ, S., BERTRAND, J.A.M., Wang, H.-Y., CHEN, W.-J., HO, H.-C., MOTOMURA, H.,  HATA, H., SADO, T., & MIYA, M. (2017) Molecular systematics of the anchovy genus Encrasicholina in the Northwest Pacific PLoS ONE, 12(7): e0181329 (link).

    16. LAVOUÉ, S., BERTRAND, J.A.M., CHEN, W.-J., HO, H.-C., MOTOMURA, H., SADO, T., & MIYA, M. (2017) Phylogenetic position of the rainbow sardine Dussumieria (Dussumieriidae) and its bearing on the early evolution of the Clupeoidei. Gene 623, 41-47 (link).

    17. LO, P.-C., LIU, S.-H.,. MOHD NOR, S.A. & CHEN, W.-J. (2017) Molecular exploration of hidden diversity in the Indo-West Pacific sciaenid clade. PLoS ONE 12(4): e0176623 (link).

    18. HIRT, M.V., ARRATIA, G.CHEN, W.-J., MAYDEN, R.L., TANG, K.L., WOOD, R.M., & SIMONS, A.M. (2017) Effects of gene choice, base composition and rate heterogeneity on inference and estimates of divergence times in cypriniform fishes. Biological Journal of the Linnean Society 121, 319-339 (link).

    19. HUNG, K.-W., RUSSELL, B.C., CHEN, W.-J. (2017) Molecular systematics of threadfin breams and relatives (Teleostei, Nemipteridae). Zoologica Scripta 46, 536-551 (link).

    20. BERTRAND, J.A.M., BORSA, P., & CHEN, W.-J. (2017) Phylogeography of the sergeants Abudefduf sexfasciatus and A. vaigiensis reveals complex introgression patterns between two widespread and sympatric Indo-West Pacific reef fishes. Molecular Ecology 26, 2527–2542 (link).

    21. FRICKE, R., CHEN, J.-N., CHEN, W.-J. (2017) New case of lateral asymmetry in fishes: a new subfamily, genus and species of deep water clingfishes from Papua New Guinea, western Pacific OceanComptes Rendus Biologies 340, 47–62 (link).


    22. CHEN, W.-J., CHEN, J.-N., PERNET, E.-J., & OLU, K.(2016) First record of the coloured righteye flounder, Poecilopsetta colorata (Teleostei: Poecilopsettidae) from the Sakalaves seamounts in the Mozambique Channel. Marine Biodiversity Records 9:83 (link).

    23. LEE, H., SAMADI, S., PUILLANDRE, N., TSAI, M-H, DAI, C.-F., CHEN, W.-J. (2016) Eight new mitogenomes for exploring the phylogeny and classification of Vetigastropoda. Journal of Molluscan Studies 82, 534-541 (link).

    24. CHEN, W.-J., HO, H.-C., & BORSA, P. (2016) Taiwanese records of oblong large-eye seabream Gymnocranius oblongus (Teleostei: Lethrinidae) and other rare or undetermined large-eye seabreams. Frontiers in Marine Science 3:107 (link).

    25. CHEN, C.-J., SHIKINA, S., CHEN, W.-J., CHUNG, Y.-J., CHIU, Y.-L., BERTRAND, J.A.M., LEE, Y-.H.,& CHANG, C.-F. (2016) A novel female-specific and sexual reproduction-associated Dmrt gene discovered in the stony coral, Euphyllia ancora. Biology of Reproduction 94(2) 40, 1-13 (link).

    26. BORSA, P., DURAND, J.-D., CHEN, W.-J., NICOLAS, H., MUTHS, D., MOU-THAM, G., KULBICKI, M (2016) Comparative phylogeography of the western Indian Ocean reef fauna. Acta Oecologica 72, 72-86 (link).


    27. CHEN, J.-N.,SAMADI, S., & CHEN, W.-J. (2015) Elopomorpha (Teleostei) as a new model fish group for evolutionary biology and comparative genomics. In: Pontarotti, P. (ed.). Evolutionary biology: Biodiversification from genotype to phenotype. Springer Verlag., Switzerland. pp. 329-344 (link).

    28. LO, P.-C., LIU, S.-H., CHAO, N.L., NUNOO, F.K.E, MOK, H.-K. CHEN, W.-J. (2015) A multi-gene dataset reveals a tropical New World origin and Early Miocene diversification of croakers (Perciformes: Sciaenidae). Molecular Phylogenetics and Evolution 88, 132–143 (link).

    29. SAMADI, S., PUILLANDRE, P., BOISSELIER, M.-C., CHEN, W.-J., CORBARI, L., MAESTRATI, P., MANA, R., PANTE, E., THUBAUT, J., ZUCCON, D. & HOURDEZ, S. (2015). Patchiness of deep-sea communities in Papua New Guinea and potential susceptibility to anthropogenic disturbances illustrated by seep organisms. Marine Ecology 36 (suppl. 1), 109-132 (link).


    30. BORSA, P., SEMBIRING, A., FAUVELOT, C. & CHEN, W.-J. (2014) Resurrection of Indian Ocean humbug damselfish, Dascyllus abudafur (Forsskal) from synonymy with its Pacific Ocean sibling, Dascyllus aruanus (L.). Comptes Rendus Biologies 337, 709–716 (link).

    31. LIU, S.-Y.V., CHANG, F.-T., BORSA, P., CHEN, W.-J. & DAI, C.-F. (2014) Phylogeography of the humbug damselfish, Dascyllus aruanus (Linnaeus, 1758): evidence of Indo-Pacific vicariance and genetic differentiation of peripheral populations. Biological Journal of the Linnean Society 113, 931–942 (link).

    32. CHANG, C.-H., LI. F. SHAO, K.-T., LIN, Y.-S., MOROSAWA, T. KIM, S., KOO, H. KIM, W., LEE, J.-S., HE, S., SMITH, C., REICHARD, M., MIYA, M., SADO, T., UEHARA, K., LAVOUÉ, S.,  CHEN, W.-J. & MAYDEN, R.L. (2014) Phylogenetic relationships of Acheilognathidae (Cypriniformes: Cyprinoidea) as revealed from evidence of both nuclear and mitochondrial gene sequence variation: Evidence for necessary taxonomic revision in the family and the identification of cryptic species. Molecular Phylogenetics and Evolution 81, 182-194 (link).

    33. CAMPBELL, M.A., LOPEZ, J.A, SATOH, T., CHEN, W.-J. & MIYA, M. (2014) Mitochondrial genomic investigation of flatfish monophyly. Gene 551, 176-182 (link).

    34. CHEN, W.-J., SANTINI, F., CARNEVALE, G., CHEN, J.-N., LIU, S.-H., LAVOUE, S. & MAYDEN, R.L. (2014) New insights on early evolution of spiny-rayed fishes (Teleostei: Acanthomorpha). Frontiers in Marine Science 1, 53 (link).

    35. CHEN, W.-J.LAVOUE, S., BEHEREGARAY, L. B. & MAYDEN, R.L. (2014) Historical biogeography of a new antitropical clade of temperate freshwater fishes. Journal of Biogeography 49, 1806–1818 (link).

    36. FRICKE, R., ALLEN, G.R., ANDRÉFOUËT, S., CHEN, W.-J., HAMEL, M.A., LABOUTE, P., MANA, R., HUI, T.H. & UYENO, D. (2014) Checklist of the marine and estuarine fishes of Madang District, Papua New Guinea, western Pacific Ocean, with 820 new records. Zootaxa 3832, 001–247 (link).

    37. CAMPBELL, M.A., CHEN, W.-J., & LOPEZ, J.A. (2014) Molecular data do not provide unambiguous support for the monophyly of flatfishes (Pleuronectiformes): a reply to Betancur-R and Ortí. Molecular Phylogenetics and Evolution 75, 149-153 (link).

    38. CHEN, J.-N., J., LOPEZ, J.A., LAVOUE, S., MIYA, M., & CHEN, W.-J. (2014) Phylogeny of the Elopomorpha (Teleostei): evidence from six nuclear and mitochondrial markers. Molecular Phylogenetics and Evolution 70, 152-161 (link).

    39. LAVOUE, S., KONSTANTINIDIS, P., & CHEN, W.-J. (2014) Progress in Clupeiform Systematics. In Biology and Ecology of Anchovies and Sardines (K. Ganias, ed). Science Publishers, Enfield, New Hampshire. pp. 1-42 (link).


    40. BORSA, P., HSIAO, D.-R., CARPENTER, K.E., & CHEN, W.-J. (2013) Cranial morphometrics and mitochondrial DNA sequences distinguish cryptic species of the longface emperor (Lethrinus olivaceus), an emblematic fish of Indo-West Pacific coral reefs. Comptes Rendus Biologies 336, 505-514 (link).

    41. CAMPBELL, M.A., CHEN, W.-J., & LOPEZ, J.A. (2013) Are flatfishes (Pleuronectiformes) monophyletic? Molecular Phylogenetics and Evolution 69664-673 (link).

    42. CHEN, W.-J., LAVOUE, S, & MAYDEN, R.L. (2013) Evolutionary origin and early biogeography of otophysan fishes (Ostariophysi: Teleostei). Evolution 67, 2218-2239 (link).

    43. HO, H.-C., & CHEN, W.-J. (2013) DNA sequences and morphological variation in Lophiodes iwamotoi Ho, Serét & Shao, 2011 based on new material from New Caledonia. Zootaxa 3682, 594-598 (link).

    44. BORSA, P., ARLYZA, I.S., CHEN, W.-J., DURAND, J.-D., MEEKAN, M.G., & SHEN, K.-N. (2013) Resurrection of New Caledonian maskray Neotrygon trigonoides (Myliobatoidei: Dasyatidae) from synonymy with N. kuhlii, based on cytochrome-oxidase I gene sequences and spotting patterns. Comptes Rendus Biologies 336, 221-232 (link). 

    45. BORSA, P., BEAREZ, P., & CHEN, W.-J. (2013) Gymnocranius superciliosus and Gymnocranius satoi, two new large-eye breams (Sparoidea: Lethrinidae) from the Coral Sea and adjacent regions. Comptes Rendus Biologies 336, 233-240 (link).

    46. LAVOUE, S, MIYA, M., MUSIKASINTHORN, P., CHEN, W.-J., &  NISHIDA, M. (2013). Evidence for an Indo-West Pacific Origin of the Clupeoidei (Teleostei: Clupeiformes). PloS One 8, e56485 (link).


    47. CHEN, W.-J., & MAYDEN, R. L. (2012) Phylogeny of suckers (Teleostei: Cypriniformes: Catostomidae): further evidence of relationships provided by the single-copy nuclear gene IRBP2. Zootaxa 3586, 195-210 (link).

    48. DURAND, J.-D., CHEN, W.-J., SEN, K.-N., ZHANG, C., & BORSA, P. (2012) Genus-level taxonomic changes imposed by the mitochondrial phylogeny of grey mullets (Teleostei: Mugilidae). Comptes Rendus Biologies 335, 687-697 (link).

    49. YANG, L., ARUNACHALAM, M., SADO, T., LEVIN, B.A., GOLUBTSOV, A.S., FREYHOF, J., FRIEL, J.P., CHEN, W.-J., HIRT, M.V., MANICKAM, R., AGNEW, M.K., SIMONS, A.M., SAITOH, K., MIYA, M., MAYDEN, R.L. & HE, S. (2012) Molecular phylogeny of the cyprinid tribe Labeonini (Teleostei: Cypriniformes). Molecular Phylogenetics and Evolution 65, 362-379 (link).

    50. DURAND, J.-D., SEN, K.-N., CHEN, W.-J., JAMANDRE, B.-W., BLEL, H. DIOP, K., NIRCHIO, M., GARCIA DE LEÓN, J.F., WHITFIELD, A.K., CHANG, C.-W. & BORSA, P. (2012) Systematics of the grey mullets (Teleostei: Mugiliformes: Mugilidae): molecular phylogenetic evidence challenges two centuries of morphology-based taxonomic studies. Molecular Phylogenetics and Evolution 64, 73-92 (link).


    51. TANG, K. L., AGNEW, M. K., CHEN, W.-J., RALEY, M. E., SADO, T., SCHNEIDER, L. M., YANG, L., BART, H. L., HE, S., LIU, H., MIYA, M., SAITOH, K., SIMONS, A. M., WOOD, R. M. & MAYDEN, R. L. (2011) Phylogeny of the gudgeons (Teleostei: Cyprinidae: Gobioninae). Molecular Phylogenetics and Evolution, 61:103-124 (link).

    52. CORIO-COSTET, M-F., DUFOUR, M-C., CIGNA, J., ABADIE, P. & CHEN, W-J. (2011) Diversity and fitness of Plasmopara viticola isolates resistant to QoI fungicides. European Journal of Plant Pathology 129:315-329 (link). 


    53. MAYDEN, R. L. & CHEN, W-J. (2010) The world's smallest vertebrate species of the Genus Paedocypris: A new family of freshwater fishes and the sister group to the world's most diverse clade of freshwater fishes (Teleostei: Cypriniformes). Molecular Phylogenetics and Evolution 57:152-175 (Link).

    54. SAITOH, K., CHEN, W-J. & MAYDEN, R.L. (2010) Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution 56(3): 1001-1010 (Link). 

    55. CHEN, W-J. & MAYDEN, R.L. (2010) A phylogenomic perspective on the new era of ichthyology. BioScience 60(6): 421-432 [view PDF].

    56. BORSA, P., COLLET, A., CARASSOU, L., PONTON, D. & CHEN, W-J. (2010) Multiple nuclear and mitochondrial genotyping identifies emperors and large-eye breams (Teleostei: Lethrinidae) from New Caledonia and reveal new large-eye bream species. Biochemical Systematics and Ecology 38: 370-389 (Link).

    57. BORSA, P., BEAREZ, P., & CHEN, W-J. (2010) Gymnocranius oblongus, a new large-eye bream species from New Caledonia (Teleostei: Lethrinidae). Comptes Rendus Biologies 333: 241-247 (Link).


    58. CHEN, W-J., LHEKNIM, V. & MAYDEN, R.L. (2009) Molecular phylogeny of the Cobitoidea (Teleostei: Cypriniformes) revisited: Position of enigmatic loach Ellopostoma Vaillant resolved with six nuclear genes. Journal of Fish Biology 75(9): 2197-2208 (Link) [view PDF].

    59. CHEN, W-J. & MAYDEN, R.L. (2009) Molecular systematics of the Cyprinoidea (Teleostei: Cypriniformes), the world's largest clade of freshwater fishes: further evidence from six nuclear genes. Molecular Phylogenetics and Evolution 52(2): 544-549 (Link) [view PDF].

    60. MAYDEN, R.L., CHEN, W-J., BART, H.L., DOOSEY, M.H., SIMONS, A.M., et al. (2009) Reconstructing the phylogenetic relationships of the Earth's most diverse clade of freshwater fishes - order Cypriniformes (Actinopterygii: Ostariophysi): a case study using multiple nuclear loci and the mitochondrial genome. Molecular Phylogenetics and Evolution 51(3): 500-514 (Link) [view PDF] (download supp. material).


    61. CORIO-COSTET, M-F., MARTINEZ, F., DELMOTTE, F.,  DOUENCE, L., RICHART-CERVERA, S. & CHEN, W-J. (2008) Resistance of Plasmopara viticola to QoIs fungicides: Origin and Diversity. In: Modern fungicides and antifungal compounds V. Dehne, H.W., Deising, H.B., Gisi, U., Kuck, K.H., Russell, P.E. & Lyr, H. (eds). DPG, Braunschweig, Germany, pp. 107-112 .

    62. CHEN, W-J., MIYA, M., SAITOH, K. & MAYDEN, R.L. (2008) Phylogenetic utility of two existing and four novel nuclear gene loci in reconstructing Tree of Life of ray-finned fishes: The order Cypriniformes (Ostariophysi) as a case study. Gene 423(2): 125-134 (Link) [view PDF] (download supp. data).

    63. SAITOH, K. & CHEN, W-J. (2008) Reducing cloning artifacts for recovery of allelic sequences by T7 endonuclease I cleavage and single re-extension of PCR products - A benchmark. Gene 423(1): 92-95 (Link) [view PDF].

    64. MAYDEN, R.L., TANG, K.L., WOOD, R.M., CHEN, W-J.et al. (2008) Inferring the Tree of Life of the order Cypriniformes, the earth’s most diverse clade of freshwater fishes: implications of varied taxon and character sampling. Journal of Systematics and Evolution 46(3): 424–438 (Link) [view PDF].

    65. CONWAY, K. W., CHEN, W-J. & MAYDEN, R. L. (2008) The 'Celestial Pearl danio' is a miniature Danio (s.s) (Ostariophysi: Cyprinidae): evidence from morphology and molecules. Zootaxa 1686: 1-28 [view PDF] (media).

    66. HE, S., GU, X., MAYDEN, R. L., CHEN, W-J., CONWAY, K. W. & CHEN, Y. (2008) Phylogenetic position of the enigmatic genus Psilorhynchus (Ostariophysi: Cypriniformes): evidence from the mitochondrial genome. Molecular Phylogenetics and Evolution 47(1): 419-425 (Link) [view PDF].

    67. HE, S., MAYDEN, R. L., WANG, X., WANG, W., TANG, K. L., CHEN, W-J. & CHEN, Y. (2008) Molecular phylogenetics of the family Cyprinidae (Actinopterygii: Cypriniformes) as evidenced by sequence variation in the first intron of S7 ribosomal protein-coding gene: further evidence from a nuclear gene of the systematic chaos in the family. Molecular Phylogenetics and Evolution 46(3): 818-829 (Link) [view PDF].


    68. CHEN, W-J.DELMOTTE, F.RICHARD-CERVERA, S., DOUENCE, L., GREIF, C. & CORIO-COSTET, M-F. (2007) At least two origins of fungicide resistance in grapevine downy mildew populations. Applied and Environmental Microbiology 73: 5162-5172 (Link) [view PDF].

    69. CHEN, W-J., RUIZ-CARUS, R. & ORTÍ, G. (2007) Relationships among four genera of mojarras (Teleostei: Perciformes: Gerreidae) from the western Atlantic and their tentative placement among percomorph fishes. Journal of Fish Biology 70 (supplement B): 202-218 (Link) [view PDF].


    70. DELMOTTE, F., CHEN, W-J., RICHARD-CERVERA, S., GREIF, C. PAPURA, D., GIRESSE, X., MONDOR-GENSON, G. & CORIO-COSTET, M-F. (2006) Microsatellite DNA markers for Plasmopara viticola, the causal agent of downy mildew of grapes. Molecular Ecology Notes 6(2):379-381 (Link) [view PDF].

     Before 2005 

    71. CHEN, W-J., ORTÍ, G. & MEYER, A. (2004) Novel evolutionary relationships among four fish model systems. Trends in Genetics 20(9): 424-431 (Link) [view PDF].

    72. LUO, J., YANG, D., SUZUKI, H., WANG, Y., CHEN, W-J., CAMPBELL, K. L. & ZHANG, Y. (2004) Molecular Phylogeny and Biogeography of Oriental Voles: Genus Eothenomys (Muridae, Mammalia). Molecular Phylogenetics and Evolution 33(2): 349-362 (Link) [view PDF].

    73. LÓPEZ, J. A., CHEN, W-J. & ORTÍ, G. (2004) Esociform phylogeny. Copeia 2004(3): 449-464 (Link) [view PDF].

    74. CHEN, W-J., BONILLO, C. & LECOINTRE, G. (2003) Repeatability of clades as a criterion of reliability: a case study for molecular phylogeny of acanthomorph (teleostei) with larger number of taxa. Molecular Phylogenetics and Evolution 26(2): 262-288 (Link) [view PDF].

    75. DEROME, N., CHEN, W-J., DETTAÏ, A., BONILLO, C. & LECOINTRE, G. (2002) Phylogeny of antarctic dragon fishes (bathydraconidae, notothenioidei, teleostei) and related families based on their anatomy and two mitochondrial genes. Molecular Phylogenetics and Evolution24(1): 139-152 (Link) [view PDF].

    76. CHEN, W-J., BONILLO, C. & LECOINTRE, G. (1998) Phylogeny of the Channichthyidae (Notothenioidei, Teleostei) based on two mitochondrial genes. In: di Prisco, G., Pisano, E. and Clarke, A. (eds.). Fishes of Antarctica. A biological Overview. Springer Verlag. pp. 287-298 [view PDF].

    N.B. Reprints (PDF) of my published papers are provided as exchange of information for personal use only. All copyrights remain with the original publishers of the articles.


    戴昌鳳 & 陳韋仁 (2018) 第十章: 海洋生物多樣性、演化和保育 in 臺灣區域海洋學 (第二版), 詹森主編, 台大出版中心.

    BORSA, P., LEMER, S., CHEN, W-J., COLLET, A., CARASSOU, L., PONTON, D. (2009) Identification par barcode (séquences nucléotidiques d’un fragment du gène du cytochrome b) des larves de Lethrinidae capturées en baies de Dumbéa et Ouano (Nouvelle-Calédonie). Rapport d'opération ZoNéCo. IRD, Nouméa, 60 pp.

    CHEN, W-J. (2001) La répétitivité des clades omme critère de la fiabilité: application à la phylogénie des Acanthomorpha (Teleostei) et des Notothenioidei (acanthomorphes antarctiques). Ph.D. thesis. University of Paris VI.

    CHEN, W-J. (1997) Phylogénie moléculaire des Channichthyidae, famille de téléostéen antarctiques. DEA thesis. University of Paris VI.


    • Ministry of Science and Technology (formerly, National Science Council), Taiwan (2009 ~ present)
    • National Science Center, Poland (2013)
    • African Coelacanth Ecosystem Programme (ACEP) of South Africa (2012)
    • National Fund for Scientific and Technological Development (FONDECYT), Chile (2010)
    • Associate editor for a specialty section of Frontiers in Marine Science - Marine Systematics and Taxonomy  (2014 ~ present)
    • Molecular Phylogenetics and Evolution (2013, 2014, 2016-2018)
    • Journal of Fish Biology (2004, 2016, 2017)
    • Zoological Studies (2017, 2018)
    • Systematic Biology (2016)
    • Zoology (2016)
    • DNA Barcodes (2015)
    • Cybium (2015)
    • Marine Biodiversity records (2015)
    • Zoological Scripta (2011, 2013, 2015)
    • Journal of Ichthyology (2014)
    • Journal of Systematics and Evolution (2012)
    • Marine Ecology Progress Series (2011)
    • Zootaxa (2010, 2019)
    • Systematics and Biodiversity (2010)
    • Biology Letter (2009)
    • Hydrobiologia (2009)
    • Ichthyological Research (2006)
    • Journal of Biogeography (2006)
    • Molecular Biology and Evolution (2003)


    • Molecular phylogenetics
    • Vertebrate zoology
    • Marine fish taxonomy: a practice work
    • Advanced course of evolutionary biology
    • Basic in theoretical evolutionary Ecology
    • Marine biotechnology: case studies
    • Methods in marine genetic data analyses
    • Evolutionary Biology and Population Genetics: Theory and Practice
    • Paper reading in marine biology (master and Ph.D. level)

    Yu-Anne & Wei-Jen 2010