*=Natural history notes, semi-popular articles, editorials, technical notes
Underline = student collaborator for which I was the primary advisor

In Press

91. Joyce, W.G., Anquetin, E.-A. Cadena, J. Claude. I.G. Danilov, S.W. Evers, G.S. Ferreira, A.D. Gentry, G.L. Georgalis, T.R. Lyson, A. Pérez-García, M. Rabi, J. Sterli, N. S. Vitek, J.F. Parham. In Press. A nomenclature for fossil and living turtles using phylogenetically defined clade names. Swiss Journal of Palaeontology

90. Türkozan, O., Ç. Karacaoğlu, J.F. Parham. In Press/2021. Reconstructions of the past distribution of Testudo graeca mitochondrial lineages in the Middle East and Transcaucasia support multiple refugia since the Last Glacial Maximum. The Herpetological Journal 31.

89. Wang, J., Y. Liu, J.J. Fong, J.F. Parham, H. Shi. In Press. Reproductive ecology of the Hainan four eye-spotted turtle (Sacalia insulensis) on Hainan Island, China. Chelonian Conservation and Biology.


88. Parham, J.F., T.J. Papenfuss, A.B. Sellas, B.L. Stuart, W.B. Simison. 2020. Genetic variation and admixture of red-eared sliders (Trachemys scripta elegans) in the USA. Molecular Phylogenetics and Evolution 145(2020):10672.

87. Biewer, J.N., J. Velez-Juarbe, J.F. Parham (corresponding author). In Press. Insights on the dental evolution of walruses based on new fossil specimens from California. Journal of Vertebrate Paleontology e1833896:1-42.

86. Simison, W.B., J.F. Parham, T.J. Papenfuss, A.W. Lam, J.B. Henderson. 2020. An annotated chromosome-level reference genome of the red-eared slider turtle (Trachemys scripta elegans). Genome Biology and Evolution 12(4):456-462. [Link to OPEN ACESSS article]

85. Gong, S., J. Wu, Y. Gao, J.J. Fong , J.F. Parham, and H. Shi. 2020. Integrating and updating wildlife conservation in China. Current Biology 30(16):R915-R919.

84. Joyce, W.G., J.F. Parham, J. Anquetin, J. Claude, I.G. Danilov IG, J.B. Iverson, B. Kear, T.R. Lyson, M. Rabi, J. Sterli. 2020. Pan-Testudines, Testudinata, Testudinesm Pan-Pleurodira, Pleurodira, Pan-Cryptodira, Cryptodira. Pp. 1041-1063. In: K. de Queiroz, P.D. Cantino, J.A. Gauthier (Eds.). Phylonyms: A Companion to the PhyloCode. CRC Press, Boca Raton.


83. Fong, J.J., B.L. Stuart, T.E. McCormack, J.F. Parham. 2019. First genetic data of the critically endangered Vietnamese pond turtle (Mauremys annamensis) from known-locality specimens. Current Herpetology 38(2):140-152. [download]

* Parham, J.F., M.S. Koo, W.B. Simison, A. Perkins, T.J. Papenfuss, E.N. Tennant. Conservation assessment of the California Legless Lizard (Anniella). California Department of Fish and Wildlife Grant Agreement Number: F15AF00458 (G1598091), 30 pp.


82. Magallanes, I., J.F. Parham (co-first author, corresponding author), G.-P. Santos, J. Velez-Juarbe. 2018. A new tuskless walrus from the Miocene of Orange County, California, with comments on the diversity and taxonomy of odobenids. PeerJ 6:e5708. [Link to OPEN ACCESS article]

81. Türkozan, O., F. Kiremit, B.R. Lavin, F. Bardakci, J.F. Parham. 2018. A comparison of the morphological and mitochondrial variation of spur-thighed tortoises, Testudo graeca, in Turkey. The Herpetological Journal 28:1-9. [download]

80. Gentry, A.D., J.F. Parham, D.J. Ehret, J.A. Ebersole. 2018. A new species of Peritresius Leidy, 1856 (Testudines: Pan-Cheloniidae) from the Late Cretaceous (Campanian) of Alabama, USA, and the occurrence of the genus within the Mississippi Embayment of North America. PLOS ONE 13(4):e0195651. [Link to OPEN ACCESS article]

79. Arakelyan, M., O. Türkozan (co-first author), N. Hezaveh, and J.F. Parham. 2018. Ecomorphology of tortoises (Testudo graeca complex) from the Araks River Valley. Russian Journal of Herpetology 25(4):245-252.


78. Kloess, P.A., and J.F. Parham. 2017. A specimen-based approach to reconstructing the late Neogene seabird communities of California. Palaeogeography, Palaeoclimatology, Palaeoecology 468:473-484. [download] [CSUF press release]

77. Barboza, M.M., J.F. Parham, G.-P. Santos, B.N. Kussman, J. Velez-Juarbe. 2017. The age of the Oso Member, Capistrano Formation, and a review of fossil crocodylians from California. PaleoBios 34:1-16. [Link to OPEN ACCESS article] [CSUF press release]

* Parham, J.F. 2017. Recent studies on the vertebrate paleontology of Orange Co., CA. Fossil Project Newsletter 4(2):11-13.


76. Pyenson, N.D., J.F. Parham, and J. Velez-Juarbe. 2016. The dilemma of trade samples and the importance of museum vouchers: caveats from a study on the extinction of Steller’s sea cow. Biology Letters 12:20150149. http://dx.doi.org/10.1098/rsbl.2015.0149. [Link to OPEN ACCESS article] [Smithsonian article]

75. Santos, G.-P., J.F. Parham, and B.L. Beatty. 2016. New data on the ontogeny and senescence of Desmostylus. Journal of Vertebrate Paleontology 36(2):e1078344. DOI:10.1080/02724634.2016.1078344. [download] [KPCC Radio interview] [OC Register] [CSUF press release]

74. Auliya, M., S. Altherr, D. Ariano-Sanchez, E.H. Baard, C. Brown, R.M. Brown, J.-C. Cantu, G. Gentile, P. Gildenhuys, E. Henningheim, J. Hintzmann, K. Kanari, M. Krvavac, M. Lettink, J. Lippert, L. Luiselli, G. Nilson, T.Q. Nguyen, V. Nijman, J.F. Parham, S.A. Pasachnik, M. Pedrono, A. Rauhaus, D. Rueda, M.-E. Sanchez, U. Schepp, M. van Schingen, N. Schneeweiss, G.H. Segniagbeto, R. Somaweera, E.Y. Sy, O. Türkozan, S. Vinke, T. Vinke, R. Vyas, S. Williamson, and T. Ziegler. 2016. Trade in live reptiles, its impact on wild populations, and the role of the European market. Biological Conservation 204:103-119. [download] [Science Daily]


73. Crawford, N.G., J.F. Parham (co-first author, corresponding author), A.B. Sellas, B.C. Faircloth, T.C. Glenn, T.J. Papenfuss, J.B. Henderson, M.H. Hansen, and W.B. Simison. 2015. A phylogenomic analysis of turtles. Molecular Phylogenetics and Evolution 83:250-257. [download] [CAS press release] [CSUF press release] [CAS Science Now] [Editor's Choice]

72. Ksepka, D.T., J.F. Parham (co-first author), J.F. Allman, M.J. Benton, M.T. Carrano, K.A. Cranston, P.C.J. Donoghue, J.J. Head, E.J. Hermsen, R.B. Irmis, W.G. Joyce, M. Kohli, K.S. Lamm, J.S.L. Patané, M.J. Phillips, N.A. Smith, N.D. Smith, M. van Tuinen, J.L. Ware, R.C.M. Warnock, and P.D. Polly. 2015. The Fossil Calibration Database — a new resource for divergence dating. Systematic Biology 64(5):853-859. [download] [CSUF press release] [Eureka Alert]

71. Parham, J.F., T.J. Papenfuss, J.R. Buskirk, G. Parra-Olea, J.Y. Chen, and W.B. Simison. 2015. Trachemys ornata or not ornata: reassessment of a taxonomic revision for Mexican Trachemys. Proceedings of the California Academy of Sciences 62(12):359-367. [download]

70. Cadena, E.A., and J.F. Parham. 2015. Oldest known marine turtle? A new protostegid from the Lower Cretaceous of Colombia. PaleoBios 32:1-32. [download]

69. Warnock, R.C.M., J.F. Parham, W.G. Joyce, T.R. Lyson, and P.C.J. Donoghue. 2015. Calibration uncertainty in molecular dating analyses: there is no substitute for the prior evaluation of time priors. Proceedings of the Royal Society B 282:20141013. [online]

68. Rhodin, A.G.J., H. Kaiser, P.P. van Dijk, W. Wüster, M. O’Shea, M. Archer, M. Auliya, L. Boitani, R. Bour, V. Clausnitzer, T. Contreras-MacBeath, B.I. Crother, J.M. Daza, C.A. Driscoll, O. Flores-Villela, J. Frazier, U. Fritz, A. Gardner, C. Gascon, A. Georges, F. Glaw, F.G. Grazziotin, C.P. Groves, G. Haszprunar, P. Havaš, J.M. Hero, M. Hoffmann, M.S. Hoogmoed, B.D. Horne, J.B. Iverson, M. Jäch, C.L. Jenkins, R.K.B. Jenkins, A.R. Kiester, J.S. Keogh, T.E. Lacher Jr., J.E. Lovich, L. Luiselli, D.L. Mahler, D. Mallon, R. Mast, R.W. Mcdiarmid, J. Measey, R.A. Mittermeier, S. Molur, V. Mossbrugger, R. Murphy, D. Naish, M. Niekisch, H. Ota, J.F. Parham, M.J. Parr, N.J. Pilcher, R.H. Pine, A.B. Rylands, J.G. Sanderson, J. Savage, W. Schleip, G.J. Scrocchi, H.B. Shaffer, E.N. Smith, R. Sprackland, S.N. Stuart, H. Vetter, L.J. Vitt, T. Waller, G. Webb, E.O. Wilson, H. Zaher, and S. Thomson (Corresponding Author). 2015. Comment on Spracklandus Hoser, 2009 (Reptilia, Serpentes, ELAPIDAE): request for confirmation of the availability of the generic name and for the nomenclatural validation of the journal in which it was published. (Case 3601; see BZN 70: 234–237; 71: 30–38, 133–135, 181–182, 252–253). Bulletin of Zoological Nomenclature 72(1):65-78. [download]

* Polly, P.D., D.T. Ksepka, and J.F. Parham. 2015. Announcing the Fossil Calibration Series and Database. Palaeontologia Electronica 18.1.1E:1-5. [online] [CSUF press release] [Eureka Alert]


67. Parham, J.F., R.A. Otero, and M.E. Suárez. 2014. A sea turtle skull from the Cretaceous of Chile with comments on the taxonomy and biogeography of Euclastes (formerly "Osteopygis"). Cretaceous Research 49:181-189. [download]

66. Pyenson, N.D., C.S. Gutstein, J.F. Parham, J.P. Le Roux, C. Carreño Chavarría, A. Metallo, V. Rossi, H. Little, A.M. Valenzuela-Toro, J. Velez-Juarbe, C.M. Santelli, D. Rubilar Rogers, M.A. Cozzuol, and M.E. Suárez. 2014. Repeated mass stranding of Miocene marine mammals from the Atacama of Chile point to sudden death at sea. Proceedings of the Royal Society B 281:20133316. [Link to OPEN ACCESS article] [Link to CERRO BALLENA WEBSITE] [ABC National News] [BBC] [LA Times] [Washington Post] [USA Today] [Time]

65. Pyenson, N.D., N.P. Kelley, and J.F. Parham. 2014. Marine tetrapod macroevolution: Physical and biological drivers on 250 million years of invasions and evolution in ocean ecosystems. Palaeogeography, Palaeoclimatology, Palaeoecology 400:1-8. [download] This article is the first of a special volume of Palaeogeography, Palaeoclimatology, Palaeoecology that includes seven original papers about "Physical drivers in the evolution of marine tetrapods" edited by Pyenson, N.D., N.P. Kelley, and J.F. Parham. The entire volume (400:1-74) can be found here: [Special Volume]

64. Rocha, L.A., A. Aleixo, G. Allen, F. Almeda, C.C. Baldwin, M.V.L. Barclay, J.M. Bates, A.M. Bauer, F. Benzoni, C.M. Berns, M.L. Berumen, D.C. Blackburn, S. Blum, F. Bolaños, R.C.K. Bowie, R. Britz, R.M. Brown, C.D. Cadena, K. Carpenter, L.M. Ceríaco, P. Chakrabarty, G. Chaves, J.H. Choat, K.D. Clements, B.B. Collette, A. Collins, J. Coyne, J. Cracraft, T. Daniel, M.R. de Carvalho, K. de Queiroz, F. Di Dario, R. Drewes, J.P. Dumbacher, A. Engilis Jr., M.V. Erdmann, W. Eschmeyer, C.R. Feldman, B.L. Fisher, J. Fjeldså, P.W. Fritsch, J. Fuchs, A. Getahun, A. Gill, M. Gomon, T. Gosliner, G.R. Graves, C.E. Griswold, R. Guralnick, K. Hartel, K.M. Helgen, H. Ho, D.T. Iskandar, T. Iwamoto, Z. Jaafar, H.F. James, D. Johnson, D. Kavanaugh, N. Knowlton, E. Lacey, H.K. Larson, P. Last, J.M. Leis, H. Lessios, J. Liebherr, M. Lowman, D.L. Mahler, V. Mamonekene, K. Matsuura, G.C. Mayer, H. Mays Jr., J. McCosker, R.W. McDiarmid, J. McGuire, M.J. Miller, R. Mooi, R.D. Mooi, C. Moritz, P. Myers, M.W. Nachman, R.A. Nussbaum, D. Ó Foighil, L.R. Parenti, J.F. Parham, E. Paul, G. Paulay, J. Pérez-Emán, A. Pérez-Matus, S. Poe, J. Pogonoski, D.L. Rabosky, J.E. Randall, J.D. Reimer, D.R. Robertson, M.-O. Rödel, M.T. Rodrigues, P. Roopnarine, L. Rüber, M.J. Ryan, F. Sheldon, G. Shinohara, A. Short, W.B. Simison, W.F. Smith-Vaniz, V.G. Springer, M. Stiassny, J.G. Tello, C.W. Thompson, T. Trnski, P. Tucker, T. Valqui, M. Vecchione, E. Verheyen, P.C. Wainwright, T.A. Wheeler, W.T. White, K. Will, J.T. Williams, G. Williams, E.O. Wilson, K. Winker, R. Winterbottom, and C.C. Witt. 2014. Specimen collection: An essential tool. Science 344(6186):814-815.


63. Parham, J.F., T.J. Papenfuss, P.P. van Dijk, B.S. Wilson, C. Marte, L. Rodriguez Schettino, and W.B. Simison. 2013. Genetic introgression and hybridization in Antillean freshwater turtles (Trachemys) revealed by coalescent analyses of mitochondrial and cloned nuclear markers. Molecular Phylogenetics and Evolution 67:176-187.

[Link to OPEN ACCESS article] [CAS press release]

62. Papenfuss, T.J., and J.F. Parham. 2013. Four new species of California legless lizards (Anniella). Breviora 536:1-7. [download] [CNN, includes VIDEO] [Los Angeles Times] [San Francisco Chronicle] [New York Times]

[National Geographic] [Scientific American] [UC Berkeley press release] [CSU Fullerton press release]

[CAS Science Today]

61. Joyce, W.G., J.F. Parham, T.R. Lyson, R.C.M. Warnock, and P.C.J. Donoghue. 2013. A divergence dating analysis of turtles using fossil calibrations: an example of best practices. Journal of Paleontology 87(4):612-634. [download]

* Simison, W.B., A.B. Sellas, K.A. Feldheim, and J.F. Parham. 2013. Isolation and characterization of microsatellite markers for identifying hybridization and genetic pollution associated with red-eared slider turtles (Trachemys scripta elegans). Conservation Genetics Resources 5(4):1139-1140. [download]


60. Parham, J.F., P.C.J. Donoghue, C.J. Bell, T.D. Calway, J.J. Head, P.A. Holroyd, J.G. Inoue, R.B. Irmis, W.G. Joyce, D.T. Ksepka, J.S.L. Patané, N.D. Smith, J.E. Tarver, M. Van Tuinen, Z. Yang, K.D. Angielczyk, J. Greenwood, C.A. Hipsley, L. Jacobs, P.J. Makovicky, J. Müller, K.T. Smith, J.M. Theodor, R.C.M. Warnock, and M.J. Benton. 2012. Best practices for justifying fossil calibrations. Systematic Biology 61(2):346-359. [Link to OPEN ACCESS article]

59. Parham, J.F., B.L. Stuart, I.G. Danilov, and N.B. Ananjeva. 2012. Genetic characterization of CITES-listed Iranian tortoises (Testudo graeca) through the sequencing of topotypic samples and a 19th century holotype. The Herpetological Journal 22:73-78. [download]

58. Otero, R.A., J.F. Parham, S. Soto-Acuña, P. Jimenez-Huidobro, and D. Rubilar-Rogers. 2012. Marine reptiles from the Late Cretaceous (early Maastrichtian) of Algarrobo, central Chile. Cretaceous Research 35:124-132. [download]

* Parham, J.F. 2012. Fossils that tell tales. DIG (June Issue):24-26.


57. Parham, J.F., and J.E. Ebersole (Eds.). Paleontology Papers in Honor of Douglas E. Jones. Alabama Museum of Natural History, Tuscaloosa, AL.

56. Bauer, A.M., J.F. Parham, R.M. Brown, B.L. Stuart, L. Grismer, T.J. Papenfuss, W. Böhme, J.M. Savage, S. Carranza, J.L. Grismer, P. Wagner, A. Schmitz, N.B. Ananjeva, and R.F. Inger. 2011. On the availability of new Bayesian-delimited gecko names and the importance of character-based species descriptions. Proceedings of the Royal Society of London Series B 278:490-492.

55. Mozaffari, O., F. Ahmadzadeh, and J.F. Parham. 2011. Eremias papenfussi sp. nov., a new lacertid lizard (Sauria: Lacertidae) from Tehran Province, Iran. Zootaxa 3114:57-62. [download]

*Ksepka, D.T., M.J. Benton, M.T. Carrano, M.A. Gandolfo, J.J. Head, E.J. Hermsen, W.G. Joyce, K.S. Lamm, J.S. Patané, M.J. Phillips, P.D. Polly, M. van Tuinen, J.L. Ware, R.C.M. Warnock, and J.F. Parham. 2011. Synthesizing and databasing fossil calibrations: divergence dating and beyond. Biology Letters 2011(7):801-803. [download]

* Parham, J.F. 2011. Book review for The Last Tortoise: A Tale of Extinction in Our Lifetime. Quarterly Review of Biology 86(3):239-240.


54. Parham, J.F., and N.D. Pyenson. 2010. New sea turtle from the Miocene of Peru and the iterative evolution of feeding ecomorphologies since the Cretaceous. Journal of Paleontology 84(2):231-247. [download]

53. Türkozan, O., F. Kiremit, J.F. Parham, K. Olgun, and E. Taskavak. 2010. A quantitative reassessment of morphology-based taxonomic schemes for Turkish tortoises. Amphibia-Reptilia 31(1):69-83. [download]

52. Papenfuss, T.J., T. Jackman, A. Bauer, B.L. Stuart, M.D. Robinson, and J.F. Parham. 2010. Phylogenetic relationships among species of Southwest Asian leaf-toed geckos (Asaccus). Proceedings of the California Academy of Sciences 61(13). [download]

51. He, B., Y. Liu, H. Shi, J. Zhang, M. Hu, Y. Ma, L. Fu, M. Hong, J. Wang, J.J. Fong, and J.F. Parham. 2010. Captive breeding of the four-eyed turtle (Sacalia quadriocellata). Asian Herpetological Research 1(2):111-117. [download]


50. Parham, J.F., and T.J. Papenfuss. 2009. Evidence for high genetic diversity among fossorial lizard populations (Anniella pulchra) in a rapidly developing landscape (Central California). Conservation Genetics 10(1):169-176. [download]

49. Papenfuss, T.J., T. Jackman, A. Bauer, B.L. Stuart, M.D. Robinson, and J.F. Parham. 2009. Phylogenetic relationships among species in the sphaerodactylid genus Pristurus. Proceedings of the California Academy of Sciences 60(19).

* Rhodin, A.G.J., J.F. Parham, P.P. van Dijk, and J.B. Iverson. 2009. Turtles of the world: Annotated checklist of taxonomy and synonymy, 2009 update, with conservation status summary. Chelonian Research Monographs 5:000.39-000.84. [download]

* Parham, J.F. 2009. Comment on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea; Reptilia, Testudines). Bulletin of Zoological Nomenclature 66(1):86. [online]


48. Parham, J.F., M. Outerbridge, B.L. Stuart, H. Erlenkeuser, D.B. Wingate, and T.J. Papenfuss. 2008. Introduced delicacy or native species? A natural Holocene origin of Bermudian terrapins revealed by fossil and genetic data. Biology Letters 4(2):216-219. [download]

* Parham, J.F. 2008. Rediscovery of an “extinct” Galápagos tortoise. Proceedings of the National Academy of the USA 105(40):15227-15228. [download]

47. Parham, J.F., and R.B. Irmis. 2008. Caveats on the use of fossil calibrations for molecular dating. American Naturalist 171(1):132-136. [download]

46. Parham, J.F. 2008. Chelonian fauna. in D. Gifford-Gonzalez. Fauna from Adrar Bous and surrounding areas. Pp. 313-353 in J.D. Clark, E.A.A. Garcea, D. Gifford-Gonzalez, A.B. Smith, and M.A.J. Williamson (Eds.; D. Gifford-Gonzalez, General Ed.) The Archaeology of a Granitic Ring Complex in Central Sahara, Niger. Archaeology, Royal Africa Museum: Tervuren, Belgium.

45. Shi, H., J.F. Parham, Z. Fan, M. Hong, and F. Yin. 2008. Evidence for the massive scale of turtle farming in China. Oryx- International Journal for Conservation 42(1):147-150. [download]

44. Danilov, I.G., and J.F. Parham. 2008. A reassessment of some poorly known turtles from the Middle Jurassic of China with comments on the antiquity of extant turtles. Journal of Vertebrate Paleontology 28(2):306-318. [download]

43. Shi, H., J.J. Fong, J.F. Parham, J. Pang, J. Wang, M. Hong, Y. Zhang. 2008. Mitochondrial variation of the "eyed" turtles (Sacalia) based on known locality and trade specimens. Molecular Phylogenetics and Evolution 49:1025-1029. [download]

42. Hong, M., H. Shi, L. Fu, S. Gong, J.J. Fong, and J.F. Parham. 2008. Scientific refutation of Traditional Chinese Medicine claims about turtles. Applied Herpetology 5(2):173-187. [download]

41. Rhodin, A.G.J., P.P. van Dijk, and J.F. Parham. 2008. Turtles of the world: Annotated checklist of taxonomy and synonymy. Chelonian Research Monographs 5:000.1-000.38. [download]

40. Arakelyan, M., and J.F. Parham. 2008. The geographic distribution of turtles in Armenia and the Nagorno-Karabakh Republic (Artsakh). Chelonian Conservation and Biology 7(1):70-77. [download]

* Arakelyan, M., J.F. Parham, O. Türkozan, and F. Danielyan. 2008. Sympatrisches Vorkommen Zweier Formen von Testudo graeca in Armenien und der Republik Nagorno -Karabakh. Marginata 2008:26-30. [in German]

* Shi, H., M. Hou, P. Pritchard, J. Peng, Z. Fan, F. Yin, T. Chen, H. Liu, J. Wang, Y. Liu (Eds.). 2008. Identification Manual for Traded Turtles in China. Contributors: P.P. van Dijk, J.F. Parham, J. Fong, B. Horne, H. Zwartpoorte, C. Schaffer, J. Tucker, B. Devaux, A. Cadi, S. Gong, G. Li. Encyclopedia of China Publishing House, Beijing, 166 pp.


39. Parham, J.F., and A.G.J. Rhodin (corresponding authors). Turtle Taxonomy Working Group. 2007. An annotated list of modern turtle terminal taxa (with comments on areas of instability and recent change). Chelonian Conservation and Biology Research Monographs 4:173-199.[download]

38. Shi H., J.F. Parham, M. Lau, and Chen T.-H. 2007. Farming endangered turtles to extinction in China. Conservation Biology 21(1):5-6. [download][This work was profiled by National Geographic]

37. Fong, J.J., J.F. Parham, Shi H., B.L. Stuart, and R.L. Carter. 2007. A genetic survey of heavily exploited, endangered turtles: Caveats on the conservation value of trade animals. Animal Conservation. 10(4):452-460. [download][This work was profiled by the Society of Conservation Biology]

36. Stuart, B.L., and J.F. Parham. 2007. Recent hybrid origin of three rare Chinese turtles. Conservation Genetics 8:169-175. [download]

35. Danilov, I.G., and J.F. Parham. 2007. The type series of ‘Sinemys’ wuerhoensis, a problematic turtle from the Early Cretaceous of China, includes at least three taxa. Palaeontology 50(2):431-444. [download]

34. Mozaffari, O., and J.F. Parham. 2007. A new species of racerunner lizard (Lacertidae: Eremias) from Iran. Proceedings of the California Academy of Sciences 58(28):569-574. [download]

33. Turtle Taxonomy Working Group. 2007. Turtle Taxonomy: methodology, recommendations, and guidelines. Chelonian Conservation and Biology Research Monographs 4:73-84. [download]


32. Parham, J.F., J.R. Macey, T.J. Papenfuss, C.R. Feldman, O. Türkozan, R. Polymeni, and J.L. Boore. 2006. The phylogeny of Mediterranean tortoises and their close relatives based on complete mitochondrial genome sequences from museum specimens. Molecular Phylogenetics and Evolution 38(1):50-64. [download]

31. Parham, J.F., C.R. Feldman, and J.L. Boore. 2006. The complete mitochondrial genome of the enigmatic bigheaded turtle (Platysternon): description of unusual genomic features and the reconciliation of phylogenetic hypotheses based on mitochondrial and nuclear DNA. BMC Evolutionary Biology 6(11):1-11. [download]

30. Parham, J.F., O. Türkozan, B.L. Stuart, M. Arakelyan, S. Shafei, J.R. Macey, and T.J. Papenfuss. 2006. Genetic evidence for premature taxonomic inflation in Middle Eastern tortoises. Proceedings of the California Academy of Sciences 57(33):955-964. [download]

29. Danilov, I.G., and J.F. Parham. 2006. A redescription of ‘Plesiochelystatsuensis, a turtle from the Late Jurassic of China, and its bearing on the antiquity of the crown clade Cryptodira. Journal of Vertebrate Paleontology 26(3):573-580. [download]

28. Danilov, I.G., and J.F. Parham (Eds.). 2006. Fossil Turtle Research Volume 1. Zoological Institute, Russian Academy of Sciences, St. Petersburg, 137 pp. [purchase]

27. Parmley, D., J.H. Hutchison, and J.F. Parham. 2006. A diverse turtle fauna from the Eocene of Georgia (USA) including the oldest records of aquatic testudinoids in southeastern North America. Journal of Herpetology 40(3):343-350. [download]

26. Lapparent de Broin, F. de, R. Bour, J.F. Parham, and J. Perälä. 2006. Eurotestudo, a new genus for the species Testudo hermanni Gmelin, 1789 (Chelonii, Testudinidae). Comptes Rendus Palevol 5:803-811. [download]

25. Macey, J.R., J.A. Schulte, J.L. Strasburg, J.A. Brisson, A. Larson, N.B. Ananjeva, Y. Wang, J.F. Parham, and T.J. Papenfuss. 2006. Assembly of the eastern North American herpetofauna: New evidence from lizards and frogs. Biology Letters 2(3):242-245. [download]

* Parham, J.F., B.L. Stuart, and N.L. Orlov. 2006. Mauremys annamensis: Distribution record for Vietnam. Herpetological Review 37(2):239. [download]

* Joyce, W.G., and J.F. Parham (Eds.). 2006. Fossil Turtle Newsletter Volume 3.


24. Parham, J.F. 2005. A reassessment of the referral of sea turtle skulls to the genus Osteopygis (Late Cretaceous, New Jersey, USA). Journal of Vertebrate Paleontology 25(1):71-77. [download]

23. Shi, H., J.F. Parham, W.B. Simison, J. Wang, S. Gong, and B. Fu. 2005. A report on the hybridization between two species of threatened Asian box turtles (Testudines: Cuora) in the wild on Hainan Island (China) with comments on the origin of ‘serrata’-like turtles. Amphibia-Reptilia 26(3):377-381. [Cover] [download]

22. Diesmos, A.C., J.F. Parham, B.L. Stuart, and R. Brown. 2005. The phylogenetic position of the recently rediscovered Philippine forest turtle (Bataguridae: Heosemys leytensis). Proceedings of the California Academy of Sciences 56(3):31-41. [download]

21. Danilov, I.G., and J.F. Parham. 2005. A reassessment of the referral of an isolated skull from the Late Cretaceous of Uzbekistan to the stem-testudinoid turtle genus Lindholmemys. Journal of Vertebrate Paleontology 25(4):784-791. [download]

20. Buskirk, J.R., J.F. Parham, and C.R. Feldman. 2005. On the hybridisation of two distantly related Asian turtles (Sacalia x Mauremys). Salamandra 41(1/2):21-26. [download]

19. Holroyd, P.A., J.F. Parham, and J.H. Hutchison. 2005. A reappraisal of some Paleogene turtles from the southeastern United States. Journal of Vertebrate Paleontology 25(4):979-982. [download]

* Parham, J.F., and S.E. Rieboldt. 2005. Contia tenuis: Reproduction. Natural history note. Herpetological Review 36(4):456.

* Joyce, W.G., and J.F. Parham (Eds.). 2005. Fossil Turtle Newsletter Volume 2.

* Shi H., D. O’Connell, J.F. Parham and K. Buley. 2005. An action plan for turtle conservation in China. Proceedings of the EAZA Conference, Kolmarden, Sweden 2004:47-57.


18. Parham, J.F., B.L. Stuart, R. Bour, and U. Fritz. 2004. Evolutionary distinctiveness of the extinct Yunnan box turtle revealed by DNA from an old museum specimen. Proceedings of the Royal Society Series B: Biology Letters 271(1556[S6]):391-394. [download]

17. Stuart, B.L., and J.F. Parham. 2004. Molecular phylogeny of the critically endangered Indochinese box turtle Cuora galbinifrons. Molecular Phylogenetics and Evolution 31:164-177. [download]

16. Joyce, W.G., J.F. Parham, and J. Gauthier. 2004. Developing a protocol for the conversion of rank-based taxon names to phylogenetically defined clade names, as exemplified by turtles. Journal of Paleontology 78(5):989-1013. [download]

15. Feldman, C.R., and J.F. Parham. 2004. Molecular systematics of Old World stripe-necked turtles (Testudines: Mauremys). Asiatic Herpetological Research 10:28-37. [download]

* Joyce, W.G., and J.F. Parham (Eds.). 2004. Fossil Turtle Newsletter Volume 1.

* Taskavak, E., M. Arakelyan, and J.F. Parham. 2004. Testudo [graeca] armeniaca Chkhikvadze & Bakradze, 1991. Manouria 7(22):7-8. [in French]

* Danilov, I.G., J.F. Parham, R, Hirayama, and G.O. Cherepanov. 2004. A report on the Symposium of Turtle Origins and Systematics held August 2003 in St. Petersburg, Russia. Herpetological Review 34(2):150. [also published in Paleontogicheskii Zhurnal]


14. Parham, J.F., and J.H. Hutchison. 2003. A new eucryptodiran turtle from the Late Cretaceous of North America (Dinosaur Provincial Park, Alberta, Canada). Journal of Vertebrate Paleontology 23(4):783-798. [download]

13. Parham, J.F., R.P. Hilton, and E.S. Göhre. 2003. The oldest turtle from California and other new records of Late Cretaceous sea turtles from the Chico Formation. PaleoBios 23(1):24-28. [download]

12. Holroyd, P.A., and J.F. Parham. 2003. The antiquity of African tortoises. Journal of Vertebrate Paleontology 23(3):688-690. [download]

11. Lynch, S.C., and J.F. Parham. 2003. First report of cheloniid sea turtles from the Miocene of California including a new species (Euclastes hutchisoni) with unusually plesiomorphic characters. PaleoBios 23(3):21-35. [download]

* Parham, J.F., and C.R. Feldman. 2003. Lampropeltis zonata: Diet. Natural history note. Herpetological Review 35(2):99-100.

* Shi H., Liu H., and J.F. Parham. 2003. The Status of Conservation, Captive Breeding, and Research of Turtles in China. China Nature 2003(1):37. [in Chinese]


10. Feldman, C.R., and J.F. Parham. 2002. A molecular phylogeny for emydine turtles: taxonomic revision and the evolution of shell kinesis. Molecular Phylogenetics and Evolution 22(3):388-398. [download]

9. Fong, J.J., J.F. Parham, and J. Fu. 2002. A reassessment of the distribution of Cuora flavomarginata Gray 1863 on Mainland China. Russian Journal of Herpetology 9(1):9-14. [download]

* Parham, J.F., and C.R. Feldman. 2002. Generic revisions of emydine turtles. Turtle and Tortoise Newsletter 6:28-30. [download]

* Parham, J.F., and T. Van Leuvan. 2002. Emys orbicularis: Distribution record for Turkey. Herpetological Review 33(2):147. [download]


8. Parham, J.F., W.B. Simison, K.H. Kozak, C.R. Feldman, and Shi, H. 2001. New Chinese turtles: Endangered or invalid? A reassessment of two species using mitochondrial DNA, allozyme electrophoresis, and known locality specimens. Animal Conservation 4(4):357-367. [download][This work was profiled by Nature]

7. Parham, J.F., and H. Shi. 2001. The discovery of Mauremys iversoni-like turtles at a turtle farm in Hainan Province, China: The counterfeit golden coin. Asiatic Herpetological Research 9:71-77. [download]

*Feldman, C.R., and J.F. Parham. 2001. Molecular systematics of emydine turtles. Chelonian Conservation and Biology 4(1):194-198. [download]

* Shi H., and J.F. Parham. 2001. Preliminary observations of a large turtle farm in Hainan Province, People's Republic of China. Turtle and Tortoise Newsletter 3:2-4. [online version]

* Parham, J.F., and Y. Wang. 2000. Palea steindachneri: Distribution record for China. Herpetological Review 31(2):111. [download]

6. Parham, J.F., and T.A. Stidham. 1999. Late Cretaceous sea turtles from the Chico Formation of California. PaleoBios 19(3):1-7. [download]

5. Parham, J.F., and D. Li. 1999. A new locality for Cuora pani Song 1984 with comments on its known range. Asiatic Herpetological Research 8:111-113. [download]

4. Parham, J.F., and G.R. Zug. 1997. Age and growth of loggerhead sea turtles (Caretta caretta) of coastal Georgia: An assessment of skeletochronological age-estimates. Bulletin of Marine Science 61(2):287-304. [download]

3. Parham, J.F., and D.E. Fastovsky. 1997. The phylogeny of cheloniid sea turtles revisited. Chelonian Conservation and Biology 2(4):548-554. [download]

2. Parham, J.F., C.K. Dodd, Jr., and G.R. Zug. 1996. Skeletochronological age estimates for the Red Hills salamander Phaeognathus hubrichti. Journal of Herpetology 30(3):401-404.

1. Zug, G.R., and J.F. Parham. 1996. Age and growth in leatherback turtles, Dermochelys coriacea (Testudines: Dermochelyidae): A skeletochronological analysis. Chelonian Conservation and Biology 2(2):244-249. [download]

* Parham, J.F., and G.R. Zug. 1996. Chelonia agassizii – valid or not? Marine Turtle Newsletter 72:2-5. [online version]