Christodoulou M.D. & Culham, A. (2021) Apples before the fall: does shape stability coincide with maturity? Quantitative Plant Biology, 2, E5. doi:10.1017/qpb.2021.5

Christodoulou M.D. & Culham, A. (2021) When do apples stop growing, and why does it matter? PLoS ONE 16(6): e0252288. doi:10.1371/journal.pone.0252288

Könyves K., Bilsborrow J., Christodoulou M.D., Culham A., & David J. (2021) Comparative plastomics of Amaryllidaceae: Inverted repeat expansion and the degradation of the ndh genes in Strumaria truncata Jacq. Accepted. PeerJ. Pre-print available at:

Christodoulou M.D., Brettschneider, J. & Steinsaltz, D. (2021) Erosion of representativeness in a cohort study. In Review. Journal of the Royal Statistical Society Series A. Pre-print available at:

Christodoulou, M. D., Clark,J, Y., Culham, A. (2020) The Cinderella discipline: morphometrics and their use in botanical classification, Botanical Journal of the Linnean Society, Volume 194, Issue 4, December 2020, Pages 385–396.

Steinsaltz, D., Christodoulou, M.D., Cohen, A.A. & Steiner, U.K. (2020) Chance Events in Aging. In: Rattan, S.I.S. (Ed.), Encyclopedia of Biomedical Gerontology. Elsevier. Vol. 1, Academic Press, pp. 386–394.

Christodoulou M.D., Battey N.H. & Culham A. (2018) Can you make morphometrics work when you know the right answer? Pick and mix approaches for apple identification. PLoS ONE 13(10): e0205357 doi: 10.1371/journal.pone.0205357

Christodoulou M.D. & Culham, A. (2015) Cyclamen rhodium subsp. peloponnesiacum near Kalavryta in June. Cyclamen: The journal of the cyclamen society. 39(2).

Working Papers

Christodoulou M.D., Kaye, E. Papavasiliou, A, & Brettschneider J. Identifying degree underperformance in tertiary education. In preparation.

Christodoulou MD, Patten H, & Steinsaltz D, Mathematical theory on biological ageing. In preparation.

Christodoulou M.D., Møllegaard, S., Brettschneider, J. A. & Steinsaltz, D. From menarche to menopause: Contrasting sequence analysis to decision theory. In preparation.

Christodoulou, M.D., Møllegaard, S., Brettschneider, J. A. & Steinsaltz, D. To impute or not to impute: Imputation protocols for Polygenic score calculation. In preparation.