Publications
Peer-reviewed research articles
Bold font identifies members of the Boyer Lab; asterisk identifies undergraduate researchers.
2024
Heine, H. L. A.*, S. Derbaraketian, R. Morisawa*, P. A. Fu*, N. H. W. Moyes*, S. L. Boyer. 2024. Machine learning approaches delimit cryptic taxa in a previously intractable species complex. In press, Molecular Phylogenetics and Evolution. (*undergraduate author) doi:10.1016/j.ympev.2024.108061
2022
Boyer, S. L., S. R. Dohr*, M. S. Tuffield*, Y. Shu*, C. D. Moore*, K. M. Hahn*, R. S. Ward*, P. Nguyen*, and R. Morisawa*. 2022. Diversity and distribution of the New Zealand endemic mite harvestman genus Aoraki (Arachnida, Opiliones, Cyphophthalmi, Pettalidae), with the description of two new species. Invertebrate Systematics 26(4): 372-387. (*undergraduate author) doi:10.1071/IS21044
Marske, K. A. and S. L. Boyer. 2022. Phylogeography reveals the complex impact of the Last Glacial Maximum on New Zealand’s terrestrial biota. Journal of the Royal Society of New Zealand. doi:10.1080/03036758.2022.2079682
2021
Tardelli Canedo, P.*, C. M. Baker, R. Morisawa*, E. J. Pessereau*, and S. L. Boyer. 2021. Phylogeography of Neopurcellia salmoni, a widespread mite harvestman from the South Island of New Zealand, with the first report of male polymorphism in the suborder Cyphophthalmi (Arachnida, Opiliones). Journal of Arachnology 49: 205–216. (*undergraduate author)
2020
Baker, C. M., S. L. Boyer, and G. Giribet. 2020. A well-resolved transcriptomic phylogeny of the mite harvestman family Pettalidae (Arachnida, Opiliones, Cyphophthalmi) reveals signatures of Gondwanan vicariance. Journal of Biogeography 47(6): 1345-1361.
2019
Fowler-Finn, K. D., S. L. Boyer, R. Ikagawa* , T. Jeffries*, P. C. Kahn*, E. M. Larsen*, D. Lee*, D. Sasson, and M. Smeester*. 2019. Qualitative and quantitative comparisons of mating behavior across multiple populations and six species of leiobunine harvestmen (Arachnida: Opiliones). Behaviour 156: 363-390. (*undergraduate coauthor)
2018
Fowler-Finn, K. D., S. L. Boyer, R. Ikagawa*, T. Jeffries*, P. C. Kahn*, E. M. Larsen*, D. Lee*, and M. Smeester*. 2018. Variation in the determinants of mating dynamics within and across five species of Leiobunum harvestmen (Arachnida: Opiliones). Biology 7(36). (Special Issue: Evolution of Mating Choice) (*undergraduate coauthor)
Kahn, P. C.*, D. C. Cao, M. M. Burns, and S. L. Boyer. 2018. Nuptial gift chemistry reveals convergent evolution correlated with antagonism in mating systems of harvestmen (Arachnida, Opiliones). Ecology and Evolution 8(14): 7103-7100. (*undergraduate author)
Oberski, J. T.*, P. P. Sharma, K. R. Jay*, M. J. Coblens*, K. A. Lemon*, J. R. Johnson*, and S. L. Boyer. 2018. A dated molecular phylogeny of mite harvestmen (Arachnida: Opiliones: Cyphophthalmi) elucidates ancient diversification dynamics in the Australian Wet Tropics. Molecular Phylogenetics and Evolution 127: 813-822. (*undergraduate author)
Sharma, P. P., C. M. Baker, J. Cosgrove, J. R. Johnson*, J. T. Oberski*, R. J. Raven. M. S. Harvey, S. L. Boyer, and G. Giribet. 2018. Testing the phylogenetic position and age of the temperate Gondwanan family Bothriuridae in the scorpion tree of life. Molecular Phylogenetics and Evolution 122: 37-45. (*undergraduate coauthor)
Sharma, P. P., J. T. Oberski, M. A. Santiago, R. Kriebel, S. M. Lipps, P. A. C. Buenavente, A. C. Diesmos, M. Janda, S. L. Boyer, R. M. Clouse, and W. C. Wheeler. 2018. There is no evidence that Podoctidae carry eggs of their own species: Reply to Machado and Wolff (2017). Molecular Phylogenetics and Evolution 129: 349-353.
2017
Sharma, P. P., M. A. Santiago, R. Kriebel, S. M. Lipps, P. A. C. Buenavente, A. C. Diesmos, M. Janda, S. L. Boyer, R. M. Clouse, and W. C. Wheeler. 2017. A multilocus phylogeny of Podoctidae (Arachnida, Opiliones, Laniatores) and parametric shape analysis reveal the disutility of subfamiliar nomenclature in armored harvestman systematics. Molecular Phylogenetics and Evolution 106: 164-173.
Shoemaker, A. K.*, M. M. Burns, and S. L. Boyer. 2017. New records of harvestmen (Arachnida, Opiliones) from Minnesota, USA. Zootaxa 4273(2): 279-286. (*undergraduate author)
2016
Boyer, S. L., T. M. Markle, C. M. Baker*, A. M. Luxbacher, and K. H. Kozak. 2016. Historical refugia have shaped biogeographical patterns of species richness and phylogenetic diversity in mite harvestmen (Arachnida, Opiliones, Cyphophthalmi) endemic to the Australian Wet Tropics. Journal of Biogeography 43(7): 1400-1411. (*undergraduate coauthor)
Clouse, R. M., P. P. Sharma, J. C. Stuart, L. R. Davis, G. Giribet, S. L. Boyer, and W. C. Wheeler. 2016. Phylogeography of the harvestman Metasiro (Arthropoda, Arachnida, Opiliones) reveals a potential solution to the Pangean paradox. Organisms Diversity and Evolution 16(1): 167-184.
Giribet, G., S. L. Boyer, C. M. Baker*, R. Fernandez, P. P. Sharma, B. L. De Bivort, S. R. Daniels, M. S. Harvey, and C. E. Griswold. 2016. A molecular phylogeny of the temperate Gondwanan family Pettalidae (Arachnida, Opiliones, Cyphophthalmi) and the limits of taxonomic sampling. Zoological Journal of the Linnean Society 178: 523-545. (*undergraduate coauthor)
Jay, K. R.*, Z. R. Popkin-Hall*, M. J. Coblens*, J. T. Oberski*, P. P. Sharma, and S. L. Boyer. 2016. New species of Austropurcellia, cryptic short-range endemic mite harvestmen (Arachnida, Opiliones, Cyphophthalmi) from Australia’s Wet Tropics biodiversity hotspot. ZooKeys 586: 37-93. (*undergraduate author)
2015
Boyer, S.L., C. M. Baker*, Z. R. Popkin-Hall*, D. I. Laukó*, H. W. Wiesner*, and R. M. Quay*. 2015. Phylogeny and biogeography of the mite harvestmen (Arachnida, Opiliones, Cyphophthalmi) of Queensland, Australia with a description of six new species from the rainforests of the Wet Tropics. Invertebrate Systematics 29: 37-70. (*undergraduate coauthor)
Hove, M. C., B. Sietman, M. Berg, E. Frost*, K. Wolf*, T. Brady*, S. L. Boyer, and D. J. Hornbach. 2015. Early life history of the sheepnose (Plethobasus cyphyus) (Mollusca: Bilvalvia: Unionoida). Journal of Natural History 50(9-10): 1-20.
Roe, K. H. and S. L. Boyer. 2015. A comparison of genetic diversity between sympatric populations of the endangered winged-mapleleaf (Quadrula fragosa) and the pimpleback (Amphinaias pustulosa) in the St. Croix River, U.S.A. American Malacological Bulletin 33(1): 1-9.
2014
Giribet, G., R. Fernández, and S. L. Boyer. 2014. On four poorly known harvestmen from New Zealand (Arachnida, Opiliones: Cyphophthalmi, Eupnoi, Dyspnoi, Laniatores). New Zealand Journal of Zoology 41(4): 223-233.
Popkin-Hall, Z. R.* and S. L. Boyer. 2014. New species of mite harvestmen from southeast Queensland, Australia greatly extend the known range of the genus Austropurcellia (Arachnida, Opiliones, Cyphophthalmi). Zootaxa 3827(4): 517-541. (*undergraduate author)
2012
Boyer, S. L. and C. N. Reuter*. 2012. New species of mite harvestmen from Australia’s Wet Tropics, with commentary on historical biogeography of the genus Austropurcellia (Opiliones, Cyphophthalmi, Pettalidae). Journal of Arachnology 40(1): 96-112. (*undergraduate coauthor)
Giribet, G., P. P. Sharma, L. R. Benavides, S. L. Boyer, R. M. Clouse, B. L. de Bivort, D. Dimitrov, G. Y. Kawauchi, J. Y. Murienne, P. J. Schwendinger. 2012. Evolutionary and biogeographic history of an ancient and global group of arachnids (Arachnida, Opiliones, Cyphophthalmi) with a new taxonomic arrangement. Biological Journal of the Linnean Society 105(1): 92-130.
Šťáhlavský, F., S. L. Boyer, M. S. Harvey, and G. Giribet. 2012. First cytogenetic study of a member of the harvestman family Pettalidae (Opiliones: Cyphophthalmi). Australian Journal of Entomology 51:299-302. pdf
Szumowski, S. C.*, S. L. Boyer, D. J. Hornbach, and M. C .Hove. 2012. Genetic diversity of two common freshwater mussels, Lampsilis cardium and Quadrula pustulosa (Bivalvia, Unionidae), in a large federally protected waterway (St. Croix River, MN/WI, USA). American Malacological Bulletin 30: 1-15. (*undergraduate author)
2011
Boyer, S. L., A. A. Howe*, N. W. Juergens*, and M. C. Hove. 2011. A DNA-barcoding approach to identifying juvenile freshwater mussels (Bivalvia: Unionidae) recovered from naturally-infested fishes. Freshwater Biology (Journal of the North American Benthological Society) 30(1): 182-194. (Special Issue: DNA Barcoding: Genomic Solutions for Benthology) (*undergraduate coauthor)
2010
Giribet, G. and S. L. Boyer. 2010. ‘Moa’s Ark’ or ‘Goodbye Gondwana’: Is New Zealand’s terrestrial invertebrate fauna ancient, recent, or both? Invertebrate Systematics 24: 1-8.
2009
Boyer, S. L. and G. Giribet. 2009. Welcome back, New Zealand: Regional biogeography and Gondwanan origin of three endemic genera of mite harvestmen. Journal of Biogeography 36 (6): 1084-1099.
2007
Boyer, S. L., J. M. Baker*, and G. Giribet. 2007. Deep genetic divergences in Aoraki denticulata (Arachnida, Opiliones, Cyphophthalmi): a widespread ‘mite harvestman’ defies DNA taxonomy. Molecular Ecology 16: 4999-5016. (*undergraduate coauthor)
Boyer, S. L., R. C. Clouse, L. Benevides, P. Schwendinger, P. P. Sharma*, I. Karunathana, and G. Giribet. 2007. Biogeography of the world: a case study from cyphophthalmid Opiliones, a globally distributed group of arachnids. Journal of Biogeography 34 (9): 2070-2085. (*undergraduate coauthor)
The paper above was featured in Science as an Editor's Choice and written up by Carl Zimmer in The New York Times as well as his textbook The Tangled Bank.
Boyer, S. L. and G. Giribet. 2007. A new model Gondwanan taxon: Systematics and biogeography of the harvestman family Pettalidae (Arachnida, Opiliones, Cyphophthalmi), with a taxonomic revision of genera from Australia and New Zealand. Cladistics 23 (4): 337-361.
2006
Giribet, G., N. Tsurusaki, and S. L. Boyer. 2006. Confirmation of the type locality and distributional range of Suzukielus sauteri (Opiliones, Cyphophthalmi) in Japan. Acta Arachnologica 55(2): 87-90.
2005
Boyer, S. L., I. Karaman, and G. Giribet. 2005. The family Sironidae (Opiliones, Cyphophthalmi) in Europe: a phylogenetic approach to Eastern Mediterranean biogeography. Molecular Phylogenetics and Evolution 36(3): 554-567.
2003
Boyer, S. L. and G. Giribet. 2003. A new Rakaia species (Opiliones, Cyphophthalmi) from Otago, New Zealand. Zootaxa 133: 1-14.
2002
Boyer, S. L., J. R. Johansen, V. R. Flechtner, and G. L. Howard. 2002. Phylogeny and genetic variance in terrestrial Microcoleus (Cyanophyceae) species based on sequence analysis of the 16S rRNA gene and associated 16S-23S ITS region. Journal of Phycology 38: 1222-1235.
Flechtner, V. R., S. L. Boyer, J. R. Johansen, and M. L. DeNoble*. 2002. Spirirestis rafaelensis gen. et sp. nov. (Cyanophyceae), a new cyanobacterial genus from arid soils. Nova Hedwigia 74: 1-24. (*undergraduate coauthor)
Giribet, G. and S. L. Boyer. 2002. A cladistic analysis of the cyphophthalmid genera (Opiliones, Cyphophthalmi). Journal of Arachnology 30: 110-128.
2001
Boyer, S. L., V. R. Flechtner, and J. R. Johansen. 2001. Is the 16S-23S internal transcribed spacer (ITS) a good tool for molecular systematics and population genetics? A case study in cyanobacteria. Molecular Biology and Evolution 18(6): 1057-1069.
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Letters, reports, and book chapters
Zeppelini, D., A. Dal Molin, C. J. E. Lamas, C. Sarmiento, C. A. Rheims, D. R. R. Fernandes, E. F. B. Lima, E. N. Silva, F. Carvalho-Filho, Ľ. Kováč, J. Montoya-Lerma, O. T. Moldovan, P. G. B. Souza-Dias, P. R. Demite, R. Feitosa, S. L. Boyer, W. M. Weiner, and W. C. Rodrigues. 2021. The dilemma of self-citation in taxonomy. Nature Ecology and Evolution 5(2): 2.
Boyer, S. L. and T. Lafrancois. 2014. Assessing the possible presence of the Baltic zooplankton Bosmina maritima in the St. Croix National Scenic Riverway. Final Report to the National Park Service.
Wolf, K., M*. C. Hove, B. E. Sietman, S. L. Boyer and D. J. Hornbach. 2012. Additional minnows and topminnow identified as suitable sheepnose (Plethobasus cyphyus) hosts. Ellipsaria 14(3): 7-8. (*undergraduate coauthor)
Hove, M. C., A. Fulton, K. M. Wolf*, B. Sietman, D. J. Hornbach, S. L. Boyer, and N. Ward. 2011. Additional suitable hosts identified for rock pocketbook (Arcidens confragosus). Ellipsaria 13(3): 6-7. (*undergraduate coauthor)
Sietman, B. E., M. C. Hove, K. Baumann*, A. Edgcumbe*, J. Jaka*, K. Linde*, J. Pfeiffer*, K. Phillips*, K. Weber*, C. Weggler*, S. L. Boyer, J. Eckstein*, N. Juergens*, and J. Ramirez*. 2010. Suitable host fishes for the Wartyback mussel, Quadrula nodulata. Ellipsaria 12: 19-20.
Sietman, B, E., K. Bloodsworth*, B. Bosman*, A. Lageer*, M. Lyons*, M. C. Hove, and S. L. Boyer. 2009. Freshwater drum confirmed as a suitable host for Leptodea, Potamilus, and Truncilla species. Ellipsaria 11(3): 18-19.
Boyer, S. L. and G. Giribet. 2007. Methods for molecular studies in systematics. Biology of the Opiliones, R. Pinto-da-Rocha, G. Machado, and G. Giribet. Harvard University Press, Cambridge, MA. 506-510.
Giribet, G. and S. L. Boyer. 2007. Pettalidae Shear ,1980. Biology of the Opiliones, R. Pinto-da-Rocha, G. Machado, and G. Giribet. Harvard University Press, Cambridge, MA. 99-101.
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In the pipeline...
Boyer, S. L., S. Derkarabetian, Z. Alafranji*, R. Morisawa*, P. Aspholm*, H. L. A. Heine*, S. Henderson*, P. A. Fu*, N. H. W. Moyes*, E. J. Pessereau*, S. A. Anderson*, and G. Giribet. Ultra-conserved elements (UCEs) resolve the phylogeny of a mite harvester genus endemic to New Zealand and contribute to identification of new species. In preparation. (*undergraduate author)
Vang, U.*, S. A. Henderson*, P. Sirvid, A. Stewart, S. L. Boyer. First report of troglobitic mite harvesters (Arachnida, Opiliones, Cyphophthalmi) from New Zealand. In preparation.
Tardelli Canedo, P., N. H. W. Moyes*, M. D. Anderson, P. Fu*, S. Derkarabetian, G. Hormiga, G. Giribet, and S. L. Boyer. Re-assessment of species-level diversity in Neopurcellia, a widespread mite harvester from the South Island of New Zealand. (*undergraduate author)
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