Publications

All my publications are available in pdf on my Research Gate webpage upon request.

Published peer-reviewed articles

2024 (3)

[29] Hendrickx, C.*, Trapman, T. H.*, Holwerda, F. M., Stein, K. H. W., Wills, S., Rauhut, O. W. M., Melzer, R. R., Van Woensel, J. and Reumer, J. W. F. 2024. A combined approach to identify isolated theropod teeth from the Cenomanian Kem Kem Group of Morocco: cladistic, discriminant, and machine learning analyses. Journal of Vertebrate Paleontology. 0 (0): e2311791. https://www.tandfonline.com/doi/full/10.1080/02724634.2024.2311791 *Authors contributed equally to this work.

[28] Hendrickx, C., Abdala, F., Filippini, F. S., Wills, S., Benson, R. and Choiniere, J. N. 2024. Evolution of postcanine complexity in Gomphodontia (Therapsida: Cynodontia). The Anatomical Record. 1–21. doi: https://doi.org/10.1002/ar.25386

[27] Delcourt, R., Brilhante, N. S., Pires-Domingues, R. A., Hendrickx, C., Grillo, O. N., Girardi, A. N., Augusta, B. G., Maciel, B. S. and Ricardo-Branco, F. 2024. Biogeography of theropod dinosaurs during the Late Cretaceous: Evidence from central South America. Zoological Journal of the Linnean Society zlad184. 1–40. https://doi.org/10.1093/zoolinnean/zlad184 

2023 (3)

[26] Abdala, F., Hendrickx, C., Jasinoski, S. C., Gaetano, L. C. and Liu, J. 2023. The pre-eminence of the Karoo Basin in the knowledge of the Permo-Jurassic cynodonts: a historical synthesis and taxonomical quantification. Palaeontologia Africana 56: 213–227. 

[25] Lei, R., Tschopp, E., Hendrickx, C., Wedel, M., Norell, M.A. and Hone, D.W.E. 2023. Bite and tooth marks on sauropod dinosaurs from the Morrison Formation. PeerJ 11:e16327: 1-34. http://doi.org/10.7717/peerj.16327    

[24] Sharma, A., Hendrickx, C. and Singh, S. 2023. First theropod record from the marine Bathonian of Jaisalmer Basin, Tethyan coast of Gondwanan India. Rivista Italiana di Paleontologia e Stratigrafia 129 (1): 49–64. 

2022 (4)

Hendrickx, C. 2022. Book review – The Dinosaurs Rediscovered. Journal of Paleontological Techniques. Book Reviews 4: 10–12.  

[23] Bell, R. P.*, Hendrickx, C.*, Pittman, M., Kaye, T. G. and Mayr, G. 2022. The exquisitely preserved integument of Psittacosaurus and the scaly skin of ceratopsian dinosaurs. Communications Biology 5 (1): 1–16.   https://www.nature.com/articles/s42003-022-03749-3 *Authors contributed equally to this work.

[22] Bell, P. R., Hendrickx, C., Pittman, M. and Kaye, T. G. 2022. Oldest preserved umbilical scar reveals dinosaurs had ‘belly buttons’. BMC Biology 20 (1): 132. https://bmcbiol.biomedcentral.com/articles/10.1186/s12915-022-01329-9   

[21] Hendrickx, C.*, Bell, R. P.*, Pittman, M., Milner, A. R. C., Cuesta, E., O’Connor, J., Loewen, M., Currie, J. P., Mateus, O., Kaye, T. and Delcourt, R., 2022. Morphology and distribution of scales and other non-feather epidermal structures in non-avialan theropod dinosaurs. Biological Reviews 97 (3): 960–1004.   https://onlinelibrary.wiley.com/doi/10.1111/brv.12829 *Authors contributed equally to this work.

2021 (4)

[20] Hendrickx, C. and Bell, P. R. 2021. The scaly skin of the abelisaurid Carnotaurus sastrei (Theropoda: Ceratosauria) from the Upper Cretaceous of Patagonia. Cretaceous Research: 104994.  https://www.sciencedirect.com/science/article/abs/pii/S0195667121002421   

[19] Tolchard, F., Kammerer, C. F., Butler, R. J., Hendrickx, C., Benoit, J., Abdala, F., and Choiniere, J. N. 2021. A new large gomphodont from the Triassic of South Africa and its implications for Gondwanan biostratigraphy. Journal of Vertebrate Paleontology: e1929265. https://www.tandfonline.com/doi/full/10.1080/02724634.2021.1929265   

[18] Meso, J.G., Hendrickx, C., Baiano, M.A., Canale, J.I., Salgado, L. and Diaz Martinez, I. 2021. Isolated theropod teeth associated with a sauropod skeleton from the Allen Formation (Campanian−Maastrichtian) of Río Negro, Argentina. Acta Palaeontologica Polonica 66 (2): 409–423. http://www.app.pan.pl/article/item/app008472020.html   

[17] Bell, P. R. and Hendrickx, C. 2021. Epidermal complexity in the theropod dinosaur Juravenator from the Late Jurassic of Germany. Palaeontology: 64 (2): 203–223. https://onlinelibrary.wiley.com/doi/abs/10.1111/pala.12517   

2020 (5)

[16] Bell, P. R. and Hendrickx, C. 2020. Crocodile-like sensory scales in a Late Jurassic theropod dinosaur. Current Biology 30 (19): R1068–R1070. https://www.sciencedirect.com/science/article/abs/pii/S0960982220312562   

[15] Hendrickx, C., Gaetano, L. C., Choiniere, J. N., Mocke, H. and Abdala, F. 2020. A new traversodontid cynodont with a peculiar postcanine dentition from the Middle/Late Triassic of Namibia and dental evolution in basal gomphodonts. Journal of Systematic Palaeontology 18 (20): 1669–1706.  https://www.tandfonline.com/doi/abs/10.1080/14772019.2020.1804470  

Tschopp, E., Araújo, R., Brusatte, S. L., Hendrickx, C., Macaluso, L., Maidment, S. C. R., Rabi, M., Rashid, D., Romano, C. and Williamson, T. 2020. Dinosaurs, But Not Only: Vertebrate Evolution in the Mesozoic. In: Martinetto, E., Tschopp, E. and Gastaldo, R. A. (eds.), Nature through Time: Virtual Field Trips through the Nature of the Past, 187–208. Springer International Publishing, Cham. https://link.springer.com/chapter/10.1007/978-3-030-35058-1_7   

[14] Hendrickx, C., Stiegler, J., Currie, P. J., Han, F., Xu, X., Choiniere, J. and Wu, X.-C. 2020. Dental anatomy of the apex predator Sinraptor dongi (Theropoda: Allosauroidea) from the Late Jurassic of China. Canadian Journal of Earth Sciences. DOI: 10.1139/cjes-2019-0231. https://www.nrcresearchpress.com/doi/abs/10.1139/cjes-2019-0231#.Xl0rBKhKjIW    

[13] Hendrickx, C., Tschopp, E. and Ezcurra, M. D. 2020. Taxonomic identification of isolated theropod teeth: the case of the shed tooth crown associated with Aerosteon (Theropoda: Megaraptora) and the dentition of Abelisauridae. Cretaceous Research: 108: 104312. https://www.sciencedirect.com/science/article/pii/S0195667118303860   

2019 (3)

[12] Hendrickx, C., Mateus, O., Araújo, R. and Choiniere, J. 2019. The distribution of dental features in non-avian theropod dinosaurs: Taxonomic potential, degree of homoplasy, and major evolutionary trends. Palaeontologia Electronica 22 (3): 1–110. https://palaeo-electronica.org/content/2019/2806-dental-features-in-theropods  

This monograph-size paper took a while to be published, with a lot of frustration, so here is my experience on the whole submission and publication process.   

[11] Hendrickx, C., Abdala, F. and Choiniere, J. N. 2019. A proposed terminology for the dentition of gomphodont cynodonts and dental morphology in Diademodontidae and Trirachodontidae. PeerJ 7: e6752.  https://peerj.com/articles/6752/  

[10] Young, C., Hendrickx, C., Challands, T.J., Foffa, D., Ross, D.A., Wilkinson, M., Panciroli, E., Butler, I., and Brusatte, S.L., 2019. New theropod dinosaur teeth from the Middle Jurassic of the Isle of Skye, Scotland. Scottish Journal of Geology. https://sjg.lyellcollection.org/content/early/2019/03/30/sjg2018-020   

2016 (3)

[9] Hendrickx, C., Abdala, F. and Choiniere, J. 2016. Postcanine microstructure in Cricodon metabolus, a Middle Triassic gomphodont cynodont from south-eastern Africa. Palaeontology: 10.1111/pala.12263. http://onlinelibrary.wiley.com/doi/10.1111/pala.12263/full  

Hendrickx, C. and Carrano, M. T. 2016. Erratum on "An overview of non-avian theropod discoveries and classification". PalArch’s Journal of Vertebrate Palaeontology 13 (2): 1–7. http://www.palarch.nl/2016/02/christophe-hendrickx-matthew-t-carrano-2016-erratum-on-an-overview-of-non-avian-theropod-discoveries-and-classification-palarchs-journal-of-vertebrate-palaeontology-13-2-2016/  

[8] Hendrickx, C., Mateus, O. and Buffetaut, E. 2016. Morphofunctional analysis of the quadrate of Spinosauridae (Dinosauria: Theropoda) and the presence of Spinosaurus and a second spinosaurine taxon in the Cenomanian of North Africa. PLoS ONE 11 (1): e0144695. http://journals.plos.org/plosone/article?id=10.1371/journal.pone.0144695#sec028  

2015 (4)

[7] Hendrickx, C., Araújo, R. and Mateus, O. 2015. The non-avian theropod quadrate I: standardized terminology with an overview of the anatomy and function. PeerJ 3:e1245; DOI 10.7717/peerj.1245. https://peerj.com/articles/1245/  

[6] Hendrickx, C., Mateus, O. and Araújo, R. 2015. A proposed terminology of theropod teeth (Dinosauria: Saurischia). Journal of Vertebrate Paleontology. 35(5):e982797. DOI: 10.1080/02724634.2015.982797.  http://www.tandfonline.com/doi/full/10.1080/02724634.2015.982797  

[5] Hendrickx, C., Hartman, S. A. and Mateus, O. 2015. An overview on non-avian theropod discoveries and classification. PalArch’s Journal of Vertebrate Palaeontology. 12 (1):1-73. http://www.palarch.nl/2015/08/christophe-hendrickx-scott-a-hartman-octavio-mateus-2015-an-overview-of-non-avian-theropod-discoveries-and-classification-palarchs-journal-of-vertebrate-palaeontology-12-1-2015/  

[4] Hendrickx, C., Mateus, O. and Araújo, R. 2015. The dentition of megalosaurid theropods. Acta Palaeontologica Polonica. 60 (3): 627-642. DOI: http://dx.doi.org/10.4202/app.00056.2013 . http://www.app.pan.pl/article/item/app000562013.html 

2014 (2)

[3] Hendrickx, C. and Mateus, O. 2014. Abelisauridae (Dinosauria: Theropoda) from the Late Jurassic of Portugal and dentition-based phylogeny as a contribution for the identification of isolated theropod teeth. Zootaxa 3759:1–74. http://dx.doi.org/10.11646/zootaxa.3759.1.1 

[2] Hendrickx, C. and Mateus, O. 2014. Torvosaurus gurneyi n. sp., the largest terrestrial predator from Europe, and a proposed terminology of the maxilla anatomy in nonavian theropods. PLoS ONE 9 (3): e88905. http://dx.doi.org/10.1371/journal.pone.0088905  

2013 (1)

[1] Araújo, R., Castanhinha, R., Martins, R. M. S., Mateus, O., Hendrickx, C., Beckmann, F., Schell, N. and Alves, L. C. 2013. Filling the gaps of dinosaur eggshell phylogeny: Late Jurassic theropod clutch with embryos from Portugal. Scientific Reports 3 (1924): 1–8. http://www.nature.com/srep/2013/130530/srep01924/full/srep01924.html  

Non peer-reviewed preprints

Bell, P. R., Hendrickx, C., Pittman, M., Kaye, T. G. and Mayr, G. 2022. The exquisitely preserved dinosaur skin of Psittacosaurus and the scaly skin of ceratopsian dinosaurs. Research Square. https://www.researchsquare.com/article/rs-1374507/v1  

Bell, P. R., Pittman, M., Kaye, T. G. and Hendrickx, C. 2020. A crocodylian-style cloaca in a non-avialan dinosaur. BioRxiv: 2020.10.11.335398. https://www.biorxiv.org/content/10.1101/2020.10.11.335398v1.full  

Hendrickx, C., Araújo, R. and Mateus, O. 2015. The nonavian theropod quadrate II: systematic usefulness, major trends and cladistic and phylogenetic morphometrics analyses. PeerJ Preprints 2:e380v1. http://dx.doi.org/10.7287/peerj.preprints.380v1  

Published oral presentations

Hendrickx, C., Abdala, F., Wills, S., Benson, R., and Choiniere, J. N., 2018. Evolution of the dentition in Gomphodontia (Therapsida: Cynodontia): Disparity, rate of evolution and dental complexity. Proceedings of the 20th Biennial Conference of the Palaeontological Society of Southern Africa, Bloemfontein, July 4 - July 6, 2018, p.18-19. 

Hendrickx, C., Mateus, O. and Araújo, R. 2014. The dentition of megalosaurids, with a proposed terminology on theropod teeth. In: Delfino, M., Carnevale, G., and Pavia, M. (Eds.). Abstract Book and Field Trip Guide, XII Annual Meeting of the European Association of Vertebrate Palaeontologists. Museo Regionale di Scienze Naturali, Regione Piemonte, Torino, p. 75. 

Hendrickx, C. and Mateus, O. 2012. Ontogenetical changes in the quadrate of basal tetanurans. In: Royo-Torres, R., Gascó, F. and Alcalá, L. (editors). 10th Annual Meeting of the European Association of Vertebrate Palaeontologists, Terruel, Spain. ¡Fundamental!, p. 101-104. 

Hendrickx, C. and Mateus, O. 2012. Deducting behaviour of dinosaurs: spinosaur theropods feeding. 9th National Congress of Ethology. (12th-13th of April 2012, Lisbon, Portugal). p. 26-27. 

Hendrickx, C. and Buffetaut, E. 2008. Functional interpretation of spinosaurid quadrates (Dinosauria: Theropoda) from the Mid Cretaceous of Morocco. In: 56th Annual Symposium of Vertebrate Palaeontology and Comparative Anatomy, Dublin, Ireland. (September 2nd-6th 2008). p. 25-26. 

Published poster presentations

Hendrickx, C., Abdala, F. and Choiniere, J. 2016. Postcanine microstructure in Cricodon metabolus, a Middle Triassic gomphodont cynodont from south-eastern Africa. 76th Annual Meeting of the Society of Vertebrate Paleontology, Salt Lake City, Utah, USA. (October 26-29, 2016). p. 152.

Hendrickx, C., Mateus, O. and Araújo, R. 2014. The distribution of dental features in non-avian theropods and a proposed terminology of theropod teeth. 74th Annual Meeting of the Society of Vertebrate Paleontology, Berlin, Germany. (November 5-8, 2014). p. 146.

Hendrickx, C., Araújo, R. and Mateus, O., 2012. The nonavian theropod quadrate : systematic usefulness, major trends and phylogenetic morphometrics analysis. 72nd Annual Meeting Society of Vertebrate Paleontology, Raleigh, USA. (October 17-20, 2012). p. 110. 

Hendrickx, C. and Mateus, O., 2011. Using dentition-based phylogeny of nonavian theropod teeth for resolving isolated teeth identification to the genus level. 55th Annual Meeting of The Palaeontological Association, Plymouth, U.K. (17th-20th December 2011). [n.b.: The abstract was not included in the abstract program. It was, however, printed in an amendment sheet.]. 

Hendrickx, C., Brusatte, S., Young, M., Rayfield, E., Ruta, M. and Barrett, P., 2009. Diversity and disparity in sauropod dinosaurs. EAVP Extraordinary Meeting, Brussels, Belgium. (February 9th-14th, 2009).

Hendrickx, C. and Buffetaut, E., 2009. Morphofunctional analysis of spinosaurid quadrates. EAVP Extraordinary Meeting, Brussels, Belgium. (February 9th-14th, 2009).

Theses

Hendrickx, C., 2015. Evolution of teeth and quadrate in non-avian Theropoda (Dinosauria: Saurischia), with the description of Torvosaurus remains from Portugal. PhD thesis. Universidade Nova de Lisboa, Portugal. 604 pp. Supervisor: Octávio Mateus. Advisers: Stephen Brusatte and Martin Sander.

Hendrickx, C., 2008. Diversity and disparity in sauropod dinosaurs. Master thesis. University of Bristol, U.K. 113 pp. Supervisors: Steve Brusatte, Mark Young, Emily Rayfield, Marcello Ruta and Paul Barrett. Adviser: Paul Upchurch.

Hendrickx, C., 2006. Analyse morphofonctionnelle des carrés de Spinosauridae. Licence thesis. Université de Liège, Belgium. 194 pp. Supervisor: Eric Buffetaut. Advisers: Edouard Poty and Jean-Marie Cordy.