A colony is a territory subject to a form of foreign rule.[1][2] Though dominated by the foreign colonizers, the rule remains separate to the original country of the colonizers, the metropolitan state (or "mother country"), which together have often been organized as colonial empires, particularly with the development of modern imperialism. This colonial administrative separation, though often blurred,[2] makes colonies neither annexed or incorporated territories nor client states. Colonies contemporarily are identified and organized as not sufficiently self-governed dependent territories. Other past colonies have become either sufficiently incorporated and self-governed, or independent, with some to a varying degree dominated by remaining colonial settler societies or neocolonialism.

While colonies often developed from trading outposts or territorial claims, such areas do not need to be a product of colonization, nor become colonially organized territories. Territories furthermore do not need to have been militarily conquered and occupied to come under colonial rule and to be considered de-facto colonies, instead neocolonial exploitation of dependency or imperialist use of power to intervene to force policy, might make a territory be considered a colony, which broadens the concept, including indirect rule or puppet states (contrasted by more independent types of client states such as vassal states). Subsequently some historians have used the term informal colony to refer to a country under a de facto control of another state. Though the broadening of the concept is often contentious.


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The word "colony" comes from the Latin word colnia, used as concept for ancient Roman outposts and eventually cities. This in turn derives from the word colnus, which was a Roman tenant farmer.

Roman colonies first appeared when the Romans conquered neighbouring Italic peoples. These were small farming settlements that appeared when the Romans had subdued an enemy in war. Though a colony could take many forms, such as a trade outpost or a military base in enemy territory, such has not been inherently colonies. Its original definition as a settlement created by people migrating from a central region to an outlying one became the modern definition.[citation needed]

Settlements that began as Roman colonia include cities from Cologne (which retains this history in its name), Belgrade to York. A tell-tale sign of a settlement within the Roman sphere of influence once being a Roman colony is a city centre with a grid pattern.[5]

The Special Committee on Decolonization maintains the United Nations list of non-self-governing territories, which identifies areas the United Nations (though not without controversy) believes are colonies. Given that dependent territories have varying degrees of autonomy and political power in the affairs of the controlling state, there is disagreement over the classification of "colony".

a group of people who leave their native country to form a settlement in a territory that their own government has claimed and forcibly taken control of: The Spanish colony in Mexico was numerous, powerful, and rich.

a number of people coming from the same country or speaking the same language, residing in a foreign country or city or in a particular section of it; enclave: There is a sizable Polish colony in Israel.

There have been several incidents of acute poisoning of honey bees covered in the popular media in recent years, but sometimes these incidents are mistakenly associated with CCD. A common element of acute pesticide poisoning of bees is, literally, a pile of dead bees outside the hive entrance. With CCD, there are very few if any dead bees near the hive. Piles of dead bees are an indication that the incident is not colony collapse disorder. Indeed, heavily diseased colonies can also exhibit large numbers of dead bees near the hive.

In October 2013, the CCD Steering Committee hosted the national stakeholder conference on honey bee health. The conference brought together a broad group of stakeholders to examine the federal government's course of action to understand colony collapse disorder and honey bee health. Based on input from the stakeholders at this conference, the CCD steering committee is drafting a revised CCD and honey bee health action plan.

Thanks and sorry that I was unclear. I want to segment the whole colony. The output I want to get is the total number of colonies and their area.

Ideally something like the image below which I have drew manually:

With your images and experimental design, finding the total area of cells should be relatively easy (as in the links provided by @melvingelbard ). However, choosing the correct input to an analysis experiment is always one of the key aspects of the experimental design, and if you want the area of each spherical colony, you have not made things easy on yourself With a relatively small number of images, you may be best of doing it manually (wand tool).

Three years later, Governor White is able to organize a relief expedition and returns to the colony. The supply ships reach Hatoraske, a harbor near Roanoke Island, and start to look for the settlers.

Wax moths arrived in the United States in 1998 in Florida. This can be a very destructive insect pest, damaging beeswax comb, comb honey, and bee-collected pollen. Wax moths are rarely the initial cause of colony failure but can overcome weak colonies.

While these four areas are easy to categorize on paper, in reality these factors often may overlap or interact with one another. Honey bees might be able to survive many of these problems if the problems occurred one at a time. But when they hit in any of a wide variety of combinations, the result can weaken and overcome the honey bee colony's ability to survive.

One problem plaguing honey bees since 2006 has been Colony Collapse Disorder (CCD), which is a syndrome specifically defined as a dead colony with no adult bees and with no dead bee bodies but with a live queen, and usually honey and immature bees, still present. CCD is not a general term that covers all managed honey bee colonies that are lost due to any reason. No scientific cause for CCD has been proven. Most research has pointed to a complex of factors being involved in the cause of CCD, and possibly not all of the same factors or the same factors in the same order are involved in all CCD incidents.

In October 2006, some beekeepers began reporting losses of 30-90 percent of their hives. While colony losses are not unexpected, especially over the winter, this magnitude of losses was unusually high.

There have also been unusual colony losses before. In 1903, in the Cache Valley in Utah, 2000 colonies were lost to an unknown "disappearing disease" after a "hard winter and a cold spring." More recently, in 1995-96, Pennsylvania beekeepers lost 53 percent of their colonies without a specific identifiable cause.

Research from ARS and other institutions has provided new management recommendations that beekeepers have begun to adopt. For example, it is now recommended that beekeepers feed honey bees more protein during times of nectar shortage such as during times of drought or in the winter. As part of this, ARS has developed a new bee diet, Megabee, now available to beekeepers. The feeding of supplemental nutrients may help to decrease winter colony losses.

ARS researchers also have been analyzing samples from healthy and CCD-struck colonies and applying a variety of stressors from the four categories of possible causes to colonies in hopes of provoking a colony response that duplicates CCD.

Studies are being conducted by ARS scientists and collaborators to look at the combined impact of two or more factors on honey bees-most recently the impact of exposure to the neonicotinoid imidacloprid and Nosema. While the dual exposure indicated some sublethal effects on individual honey bees, the overall health of the colony did not show an adverse effect.

A honey bee colony typically consists of three kinds of adult bees: workers, drones, and a queen. Several thousand worker bees cooperate in nest building, food collection, and brood rearing. Each member has a definite task to perform, related to its adult age. But surviving and reproducing take the combined efforts of the entire colony. Individual bees (workers, drones, and queens) cannot survive without the support of the colony.

Each colony has only one queen, except during and a varying period following swarming preparations or supersedure. Because she is the only sexually developed female, her primary function is reproduction. She produces both fertilized and unfertilized eggs. Queens lay the greatest number of eggs in the spring and early summer. During peak production, queens may lay up to 1,500 eggs per day. They gradually cease laying eggs in early October and produce few or no eggs until early next spring (January). One queen may produce up to 250,000 eggs per year and possibly more than a million in her lifetime.

New (virgin) queens develop from fertilized eggs or from young worker larvae not more than 3 days old. New queens are raised under three different circumstances: emergency, supersedure, or swarming. When an old queen is accidentally killed, lost, or removed, the bees select younger worker larvae to produce emergency queens. These queens are raised in worker cells modified to hang vertically on the comb surface (Figure 2). When an older queen begins to fail (decreased production of queen substance), the colony prepares to raise a new queen. Queens produced as a result of supersedure are usually better than emergency queens since they receive larger quantities of food (royal jelly) during development. Like emergency queen cells, supersedure queen cells typically are raised on the comb surface. In comparison, queen cells produced in preparation for swarming are found along the bottom margins of the frames or in gaps in the beeswax combs within the brood area. 006ab0faaa

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