A dolphin is an aquatic mammal within the infraorder Cetacea. Dolphin species belong to the families Delphinidae (the oceanic dolphins), Platanistidae (the Indian river dolphins), Iniidae (the New World river dolphins), Pontoporiidae (the brackish dolphins), and possibly extinct Lipotidae (baiji or Chinese river dolphin). There are 40 extant species named as dolphins.

Dolphins range in size from the 1.7-metre-long (5 ft 7 in) and 50-kilogram (110-pound) Maui's dolphin to the 9.5 m (31 ft) and 10-tonne (11-short-ton) orca. Various species of dolphins exhibit sexual dimorphism where the males are larger than females. They have streamlined bodies and two limbs that are modified into flippers. Though not quite as flexible as seals, they are faster; some dolphins can briefly travel at speeds of 29 kilometres per hour (18 mph) or leap about 9 metres (30 ft).[1] Dolphins use their conical teeth to capture fast-moving prey. They have well-developed hearing which is adapted for both air and water. It is so well developed that some can survive even if they are blind. Some species are well adapted for diving to great depths. They have a layer of fat, or blubber, under the skin to keep warm in the cold water.


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Dolphins are widespread. Most species prefer the warm waters of the tropic zones, but some, such as the right whale dolphin, prefer colder climates. Dolphins feed largely on fish and squid, but a few, such as the orca, feed on large mammals such as seals. Male dolphins typically mate with multiple females every year, but females only mate every two to three years. Calves are typically born in the spring and summer months and females bear all the responsibility for raising them. Mothers of some species fast and nurse their young for a relatively long period of time. Dolphins produce a variety of vocalizations, usually in the form of clicks and whistles.

Dolphins are sometimes hunted in places such as Japan, in an activity known as dolphin drive hunting. Besides drive hunting, they also face threats from bycatch, habitat loss, and marine pollution. Dolphins have been depicted in various cultures worldwide. Dolphins are sometimes kept in captivity and trained to perform tricks. The most common dolphin species in captivity is the bottlenose dolphin, while there are around 60 orcas in captivity.

The term 'dolphin' can be used to refer to most species in the family Delphinidae (oceanic dolphins) and the river dolphin families Iniidae (South American river dolphins), Pontoporiidae (La Plata dolphin), Lipotidae (Yangtze river dolphin) and Platanistidae (Ganges river dolphin and Indus river dolphin).[6][7]This term has often been applied in the US, mainly in the fishing industry, to all small cetaceans (dolphins and porpoises) are considered to be porpoises,[8] while the fish dorado is called dolphin fish.[9] In common usage the term 'whale' is used only for the larger cetacean species,[10] while the smaller ones with a beaked or longer nose are considered 'dolphins'.[11] The name 'dolphin' is used casually as a synonym for bottlenose dolphin, the most common and familiar species of dolphin.[12] There are six species of dolphins commonly thought of as whales, collectively known as blackfish: the orca, the melon-headed whale, the pygmy killer whale, the false killer whale, and the two species of pilot whales, all of which are classified under the family Delphinidae and qualify as dolphins.[13] Although the terms 'dolphin' and 'porpoise' are sometimes used interchangeably, 'porpoise' usually refers to the Phocoenidae family, which have a shorter beak and spade-shaped teeth and differ in their behavior.[12]

In 1933, three hybrid dolphins beached off the Irish coast; they were hybrids between Risso's and bottlenose dolphins.[15] This mating was later repeated in captivity, producing a hybrid calf. In captivity, a bottlenose and a rough-toothed dolphin produced hybrid offspring.[16] A common-bottlenose hybrid lives at SeaWorld California.[17] Other dolphin hybrids live in captivity around the world or have been reported in the wild, such as a bottlenose-Atlantic spotted hybrid.[18] The best known hybrid is the wholphin, a false killer whale-bottlenose dolphin hybrid. The wolphin is a fertile hybrid. Two wolphins currently live at the Sea Life Park in Hawaii; the first was born in 1985 from a male false killer whale and a female bottlenose. Wolphins have also been observed in the wild.[19]

Archaeoceti is a parvorder comprising ancient whales. These ancient whales are the predecessors of modern whales, stretching back to their first ancestor that spent their lives near (rarely in) the water. Likewise, the archaeocetes can be anywhere from near fully terrestrial, to semi-aquatic to fully aquatic, but what defines an archaeocete is the presence of visible legs or asymmetrical teeth.[23][24][25][26] Their features became adapted for living in the marine environment. Major anatomical changes include the hearing set-up that channeled vibrations from the jaw to the earbone which occurred with Ambulocetus 49 million years ago, a streamlining of the body and the growth of flukes on the tail which occurred around 43 million years ago with Protocetus, the migration of the nasal openings toward the top of the cranium and the modification of the forelimbs into flippers which occurred with Basilosaurus 35 million years ago, and the shrinking and eventual disappearance of the hind limbs which took place with the first odontocetes and mysticetes 34 million years ago.[27][28][29] The modern dolphin skeleton has two small, rod-shaped pelvic bones thought to be vestigial hind limbs. In October 2006, an unusual bottlenose dolphin was captured in Japan; it had small fins on each side of its genital slit, which scientists believe to be an unusually pronounced development of these vestigial hind limbs.[30]

Dolphins have torpedo-shaped bodies with generally non-flexible necks, limbs modified into flippers, a tail fin, and bulbous heads. Dolphin skulls have small eye orbits, long snouts, and eyes placed on the sides of its head; they lack external ear flaps. Dolphins range in size from the 1.7 m (5 ft 7 in) long and 50 kg (110 lb) Maui's dolphin to the 9.5 m (31 ft 2 in) and 10 t (11 short tons) orca. Overall, they tend to be dwarfed by other Cetartiodactyls. Several species have female-biased sexual dimorphism, with the females being larger than the males.[34][35]

All dolphins have a thick layer of blubber, thickness varying on climate. This blubber can help with buoyancy, protection to some extent as predators would have a hard time getting through a thick layer of fat, and energy for leaner times; the primary usage for blubber is insulation from the harsh climate. Calves, generally, are born with a thin layer of blubber, which develops at different paces depending on the habitat.[35][37]

Blubber is found within the dermis and subcutis layer. The dermis blends gradually with the adipose layer, which is known as fat, because the fat may extend up to the epidermis border and collagen fiber bundles extend throughout the whole subcutaneous blubber which is fat found under the skin.[43] The thickness of the subcutaneous blubber or fat depends on the dolphin's health, development, location, reproductive state, and how well it feeds. This fat is thickest on the dolphin's back and belly. Most of the dolphin's body fat is accumulated in a thick layer of blubber. Blubber differs from fat in that, in addition to fat cells, it contains a fibrous network of connective tissue.[45]

The blubber functions to streamline the body and to form specialized locomotor structures such as the dorsal fin, propulsive fluke blades and caudal keels.[43] There are many nerve endings that resemble small, onion-like configurations that are present in the superficial portion of the dermis. Mechanoreceptors are found within the interlocks of the epidermis with dermal ridges. There are nerve fibers in the dermis that extend to the epidermis. These nerve endings are known to be highly proprioceptive, which explains sensory perception.[43] Proprioception, which is also known as kinesthesia, is the body's ability to sense its location, movements and actions. Dolphins are sensitive to vibrations and small pressure changes.[46]Blood vessels and nerve endings can be found within the dermis. There is a plexus of parallel running arteries and veins in the dorsal fin, fluke, and flippers.[43] The blubber manipulates the blood vessels to help the dolphin stay warm. When the temperature drops, the blubber constricts the blood vessels to reduce blood flow in the dolphin.[47] This allows the dolphin to spend less energy heating its own body, ultimately keeping the animal warmer without burning energy as quick. In order to release heat, the heat must pass the blubber layer. There are thermal windows that lack blubber, are not fully insulated and are somewhat thin and highly vascularized, including the dorsal fin, flukes, and flippers.[48] These thermal windows are a good way for dolphins to get rid of excess heat if overheating. Additionally in order to conserve heat, dolphins use countercurrent heat exchange. Blood flows in different directions in order for heat to transfer across membranes. Heat from warm blood leaving the heart will heat up the cold blood that is headed back to the heart from the extremities, meaning that the heart always has warm blood and it decreases the heat lost to the water in those thermal windows.[48]

Some dolphins are adapted for diving to great depths. In addition to their streamlined bodies, some can selectively slow their heart rate to conserve oxygen.[52][53] Some can also re-route blood from tissue tolerant of water pressure to the heart, brain and other organs. Their hemoglobin and myoglobin store oxygen in body tissues, and they have twice as much myoglobin as hemoglobin.[54]

A dolphin ear has specific adaptations to the marine environment. In humans, the middle ear works as an impedance equalizer between the outside air's low impedance and the cochlear fluid's high impedance. In dolphins, and other marine mammals, there is no great difference between the outer and inner environments. Instead of sound passing through the outer ear to the middle ear, dolphins receive sound through the throat, from which it passes through a low-impedance fat-filled cavity to the inner ear. The ear is acoustically isolated from the skull by air-filled sinus pockets, which allow for greater directional hearing underwater.[55] Dolphins send out high frequency clicks from an organ known as a melon. This melon consists of fat, and the skull of any such creature containing a melon will have a large depression. This allows dolphins to use echolocation for orientation.[35][56][57][58][59] Though most dolphins do not have hair, they do have hair follicles that may perform some sensory function.[60] Beyond locating an object, echolocation also provides the animal with an idea on an object's shape and size, though how exactly this works is not yet understood.[61] The small hairs on the rostrum of the boto (river dolphins of South America) are believed to function as a tactile sense, possibly to compensate for the boto's poor eyesight.[62] ff782bc1db

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