It was this exact reason that I bought my next pair of Allbirds in the Tree material after reading reviews of the tree skippers. I have worn them on tropical days in England (yes, they do exist!) where my feet remained cool and bone dry after a long walk. They really are breathable!

Skippers are a group of butterflies placed in the family Hesperiidae within the order Lepidoptera (moths and butterflies). They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy places the family in the superfamily Papilionoidea, the butterflies. They are named for their quick, darting flight habits. Most have their antenna tips modified into narrow, hook-like projections. Moreover, skippers mostly have an absence of wing-coupling structure available in most moths.[1] More than 3500 species of skippers are recognized, and they occur worldwide, but with the greatest diversity in the Neotropical regions of Central and South America.[2]


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Collectively, these three groups of butterflies share many characteristics, especially in the egg, larval, and pupal stages.[2] Nevertheless, skippers have the antennae clubs hooked backward like a crochet hook, while the typical butterflies have club-like tips to their antennae, and moth-butterflies have feathered or pectinate (comb-shaped) antennae similar to moths. Skippers also have generally stockier bodies and larger compound eyes than the other two groups, with stronger wing muscles in the plump thorax, in this resembling many moths more than the other two butterfly lineages do. Unlike, for example, the Arctiinae, though, their wings are usually small in proportion to their bodies. Some have larger wings, but only rarely as large in proportion to the body as in other butterflies. When at rest, skippers keep their wings usually angled upwards or spread out, and only rarely fold them up completely.[2]

The wings are usually well-rounded with more or less sharply tipped forewings. Some have prominent hindwing tails, and others have more angled wings; the skippers' basic wing shapes vary not much by comparison to the Papilionoidea, though. Most have a fairly drab coloration of browns and greys; some are more boldly black-and-white. Yellow, red, and blue hues are less often found, but some largely brown species are quite richly colored, too. Green colors and metallic iridescence are generally absent. Sexual dichromatism is present in some; males may have a blackish streak or patch of scent scales on their forewings.[2]


Many species of skippers look very alike. For example, some species in the genera Amblyscirtes, Erynnis (duskywings), and Hesperia (branded skippers) cannot currently be distinguished in the field even by experts. The only reliable method of telling them apart involves dissection and microscopic examination of the genitalia, which have characteristic structures that prevent mating except between conspecifics.[2]

The Hesperiidae are members of the Superfamily Hesperioidea. Worldwide in distribution, skippers are richest in the tropics. More than 3,500 species are described, with approximately 275 in North America, many of which are found only in Arizona and Texas. Most skippers are small to medium, usually orange, brown, black, white, or gray. A few have iridescent colors. Skippers have large eyes, short antennae (often with hooked clubs), stout bodies, and three pairs of walking legs. Their flight is often rapid, making wing movement appear blurred. Adults of most species have long probicscises and feed on floral nectar, but some also take up nutrients from bird droppings. Males have scent scales found in modified forewing patches. Males of most species locate mates by perching (grass and giant-skippers), though some patrol, especially in the open-winged skippers. Globular eggs are laid singly.

Many Missouri species are found in locally abundant places or scattered localities near their favorite food plants. Since some food plants are located mainly in native tallgrass prairies, the skippers that feed on them form colonies in those limited habitats. Other species, which feed on a wide variety of plants, are more widespread.

The larvae of grass skippers typically feed on grasses and sedges, while the larvae of spread-winged skippers typically feed on forbs (broad-leaved wildflowers and other herbaceous plants, such as legumes or mallows) or on the leaves of woody plants such as oaks.

Many skippers have multiple broods, while some have only one generation a year. Like other butterflies, skippers hatch from eggs as caterpillars. The caterpillars grow and molt a number of times. When fully grown, the caterpillar pupates and emerges as a winged adult. Female skippers typically lay eggs singly on or near suitable host plants. Grass skipper caterpillars build shelters by rolling blades of grass around their bodies; this is how they spend the winter.

In some species, male skippers stake out a territory, perching on a prominent object and waiting for females to fly near. In other species, males seek females by patrolling an area. Male grass skippers are often seen fluttering rapidly around females, wafting the pheromones from the stigma patches on their wings, in their attempts to woo potential mates. Males also often dash out to inspect and drive away potential intruders.

Some skippers, though they may breed in our state during the summer, cannot survive Missouri winters. The sachem and fiery skipper are examples. They must recolonize our state each year, as populations that survive winters to our south gradually expand their ranges northward over the spring and summer.

We associate skippers with carefree summer days . . . sun-kissed grasslands full of wildflowers . . . gardens bright with zinnias, sunflowers, and marigolds . . . and lazy walks on dusty country roads, where flowers bloom freely in fencerow thickets.

Many predators eat skippers, including birds, mammals, reptiles, and amphibians. A skipper that falls into a lake may be eaten by a fish. But hiding in the centers of the flowers they visit are crab spiders, ambush bugs, praying mantises, and other predators that are just as hungry.

The byssus, Ottoe, and arogos skippers have localized populations limited to high-quality native prairie habitats. That several of our skippers are strongly associated with native prairies is a reminder that prairie conservation is important not just for us but for other organisms, too.

In the southeastern Cascade Mountains, mardon skippers are found in meadows in an otherwise forested landscape; a variety of grasses and sedges are used for egg-laying (and larval hosts) and females select for large, well developed plants. The historical and ongoing loss of montane meadow habitat is well-documented.

These skippers complete a single life cycle annually (univoltine).They are sedentary butterflies and do not migrate; instead, the species inhabits sites year-round (as egg, larva, pupa and adult), typically moving within only a few hundred meters of their natal locations.

Male skippers seek mates by perching on low vegetation and then darting out to inspect passing butterflies. Males that detect females commence courtship behavior; when males detect another male they engage in a territory defense behavior of tight, upward spiraling flight.

Mardon skippers primarily occur in a few small, isolated populations. Overall, their population is highly disjunct. Their range includes South Puget Sound and southeast Cascades in Washington, southwestern Oregon, and northwest California.

Climate sensitivity of this species is influenced by increased temperature, changes in precipitation patterns, early snowmelt, reduced snowpack, and increased wildfire. Population numbers vary annually in response to variable weather because Mardon skippers exhibit physiological and indirect (i.e., habitat) sensitivity to temperature and precipitation. Temperature influences butterfly behavior (e.g., mating, foraging, egg-laying time), adult life span, and larval development. Cool, wet spring conditions limit adult activity and therefore fecundity. Warming temperature may also affect phenological timing between Mardon skipper and key plant species (host and nectar plants) possibly causing low availability of nectar resources and pre-mature desiccation of larval forage, leading to reduced fitness or starvation of adults or larvae. A majority of Mardon skipper populations in WA are located in higher elevation sites, where warming temperatures lead to reduced snowpack/earlier snowmelt creating highly exposed, novel conditions for overwintering likely increasing mortality, and contribute to habitat degradation by hastening conifer encroachment in the butterfly's open, grassland habitat. Mardon skippers are also vulnerable to fire. Fire helps maintain open grassland habitat used by the Mardon skipper by preventing conifer encroachment, but Mardon skippers are highly sedentary and unable to escape fire, therefore fire can cause direct mortality of all life stages. Increasing fire frequencies may expand overall habitat area available for Mardon skipper, but could contribute to population extirpation if fire occurs in current habitat areas. 17dc91bb1f

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