Sam Simkin, Ph.D.

Staff Scientist / Quantitative Ecology
1685 38th Street, Suite 100
Boulder, CO 80301
Phone and E-mail contact info:
(720) 514-9901   
E-mail: ssimkin at                                 

Postdoc, University of Colorado, INSTAAR and Department of Ecology and Evolutionary Biology  
Ph.D., 2012, Cornell University and Cary Institute of Ecosystem Studies
B.A., 1995, Earlham College
Research Interests
• Wetland plant diversity as influenced by sulfide biogeochemistry

• Clark C.M., S.M. Simkin, E.B. Allen, W.D. Bowman, J. Belnap, M.L. Brooks, S.L. Collins, L.H. Geiser, F.S. Gilliam, S.E. Jovan, L.H. Pardo, B.K. Schulz, C.J. Stevens, K.N. Suding, H.L. Throop, and D.M. Waller. 2019. Potential vulnerability of 348 herbaceous species to atmospheric deposition of nitrogen and sulfur in the United States. Nature Plants 5:697-705. DOI:
• Perring, M.P., M. Diekmann, G. Midolo, D. Schellenberger Costa, M. Bernhardt-Römermann, J.C.J. Otto, F.S. Gilliam, P. Hedwall, A. Nordin, T. Dirnböck, S.M. Simkin, F. Máliš, H. Blondeel, J. Brunet, M. Chudomelová, T. Durak, P. De Frenne, R. Hédl, M. Kopecký, D. Landuyt, D. Li, P. Manning, P. Petrík, K. Reczynska, W. Schmidt, T. Standovár, K. Swierkosz, O. Vild, D.M. Waller, and K. Verheyen. 2018. Understanding context dependency in the response of forest understorey plant communities to nitrogen deposition. Environmental Pollution 242:1787-1799.
• Simkin, S.M., E.B. Allen, W.D. Bowman, C.M. Clark, J. Belnap, M.L. Brooks, B.S. Cade, S.L. Collins, L.H. Geiser, F.S. Gilliam, S.E. Jovan, L.H. Pardo, B.K. Schulz, C.J. Stevens, K.N. Suding, H.L. Throop, and D.M. Waller. 2016. Conditional vulnerability of plant diversity to atmospheric nitrogen deposition across the United States. Proceedings of the National Academy of Sciences of the United States of America 113:4086-4091.
• Simkin, S.M., B.L. Bedford, and K.C. Weathers. 2013. Phytotoxic sulfide more important than nutrients for plants within a groundwater-fed wetland. Ecosystems 16:1118-1129.
• Weathers, K.C., S.M. Simkin, G.M. Lovett, and S.E. Lindberg. 2006.
Empirical modeling of atmospheric deposition in mountainous landscapes. Ecological Applications 16:1590-1607.
• Simkin, S.M. and W.K. Michener. 2005.
Faunal soil disturbance regime of a longleaf pine ecosystem. Southeastern Naturalist 4:133-152.
• Driese, K.L., W.A. Reiners, G.M. Lovett, and S.M. Simkin. 2004.
A vegetation map for the Catskill Park, NY, derived from multi-temporal Landsat imagery and GIS data. Northeastern Naturalist 11:421-442.
• Simkin, S.M., D.N. Lewis, K.C. Weathers, G.M. Lovett, and K. Schwarz. 2004.
Determination of sulfate, nitrate, and chloride in throughfall using ion-exchange resins. Water, Air, & Soil Pollution 153:343-354.
• Simkin, S.M., W.K. Michener, and R. Wyatt. 2004.
Mound microclimate, nutrients and seedling survival. American Midland Naturalist 152:12-24.
• Simkin, S.M., W.K. Michener, and R. Wyatt. 2001.
Plant response following soil disturbance in a longleaf pine ecosystem. Journal of the Torrey Botanical Society 128:208-218.