427.4–423.0 Ma
Ludfordian: 425.6±0.9–423.0±2.3 Ma
Gorstian: 427.4±0.5–425.6±0.9 Ma
There is a marked increase in the plant structure complexity by the late Silurian (Ludlow)
Record of tracheids, and therefore evidence of vascular plants, in the fossil record (Edwards & Davies, 1976)
C-type tracheids were also discovered earlier in the Wenlock within Cooksonia pertonii †
Cooksonia cambrensis † and C. pertonii † are found during this time from fossils in Ireland (Edwards et al., 1979; Edwards & Richardson, 2004)
Steganotheca striata † from Wales (Edwards & Rogerson, 1979)
Baragwanathia longifolia †, Salopella australis †, Hedeia † have been reported from Australia (Tims & Chambers, 1984)
Baragwanathia † represents the earliest known vascular plant leaves (microphylls)
Bathurstia †, Distichophytum †, Hostinella †, Macivera gracilis †, and Zosterophyllum † (Kotyk et al., 2002)