The Sakmarian (293.5–290.1 Ma) is the second stage of the Early Permian (Cisuralian Epoch), occurring before the Artinskian Age, and after the Asselian Age
295.0±0.18–290.1±0.26 Ma
Phanerozoic; Mesozoic; Permian; Cisuralian (Early Permian)
Flora, spore-bearing
Forests of scale trees, ferns, horsetails, progymnosperms, and seed ferns
The dominant trees are noeggerathian progymnosperms with an emergent layer of scale trees, such as Sigillaria in some areas
Extant clubmoss groups, such as the Lycopodiales and Selaginellales, are found in the understory of some forests
There are abundant marattialean tree ferns, as well as smaller zygopterid ferns
Sphenophyllophytes are found in the understory and horsetails along rivers and wetlands
One of the earliest known true ferns, Oligosporangiopteris zhongxiangii, was found in Inner Mongolia, North China (Frojdová et al., 2021)
Flora, seed-bearing
The Cordaitales diversify in Euramerica, and the Glossopteridales diversify in Gondwana
Cycads may have been present, but there is debate on the earliest known member of this group
Giganopterids were among the most striking and important plants of the Cathaysian flora of Sino-Malaya (Wang, 1999)
Seed ferns, such as the medullosids, callistophytes, and peltasperms, were still on the landscape, surviving from the Carboniferous
Medullosids had a (semi-)self-supporting growth habit with a high water-conducting potential of these plants, thriving under a seasonally-dry climate on wet, rocky soils showing proximity to the groundwater level (Luthardt et al., 2021)