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Pragian Age (413–410.6 Ma) is the second stage of the Early Devonian Epoch that occurs after the Lochkovian Age, and before the Emsian Age. It is also known as the Siegenian in North America.
413.02–410.62 Ma
Above: A reconstruction of the Rhynie biota during the Early Devonian. Credit: Victor Leshyk
The exquisitely preserved Rhynie Chert from Scotland, dating to 407 Ma
The environment at this locality fossilized exquisite details from the Early Devonian geyser-adjacent environment
The chert was formed when silica-rich water from volcanic springs rose rapidly and petrified the early terrestrial ecosystem, in situ and almost instantaneously
Bryophyta: Sporogonites †; Riccardiothallus †
Others: Amplectosporangium †, Catenalis †, Chakassiophyton †, Changwuia †, Dawsonites †, Hostinella †, Margophyton †, Minusia †, Pilophytites †, Sciadocillus †, Stachyophyton †, Wahnbachella †
Rhyniophytes and Cooksonioids such as Celatheca †, Cooksonia †, Eogaspesia †, Hedeia †, Hsüa †, Rhynia †, Salopella †, Sartilmania †, Sciadophyton †, Sennicaulis †, Stockmansella †, Uskiella †, Yarravia †
Zosterophylls such as Bathurstia †, Deheubarthia †, Demersatheca †, Discalis †, Distichophytum †, Gosslingia †, Guangnania †, Gumuia †, Hicklingia †, Huia †, Nothia †, Oricilla †, Ramoferis †, Sawdonia †, Taeniocrada †, Tarella †, Trichopherophyton †, Thrinkophyton †, Ventarura †, Wenshania †, Yunia †, Zosterophyllum †
Lycopods such as Asteroxylon †, Baragwanathia †, Cervicornus †, Drepanophycus †, Estinnophyton †, Halleophyton †, Hueberia †, and Zhenglia †
Barinophytes, including Adoketophyton †, ?Bracteophyton †, ?Dibracophyton †, Krithodeophyton †, Pectinophyton †, and Protobarinophyton †
Basal euphyllophytes are also found in other (non-Rhynie) localities such as Psilophyton † and Polythecophyton †
Armoricaphyton chateaupannense † from the latest Pragian & Emsian of Montjean-sur-Loire, France (Gerrienne and Gensel, 2016)
Pauthecophyton gracile † from the Posongchong flora of Yunnan, China (Xue et al., 2012)
Eophyllophyton bellum †, from the Posongchong flora of Yunnan, China, is one of the first euphyllophytes with flattened appendages that appear to be small leaves (Hao, 1988; Hao & Beck, 1993)
Enstinnophyton yunnanense †, from the Posongchong flora of Yunnan, China, is a potential early equisetophyte
History and contemporary significance of the Rhynie cherts (Edwards et al., 2017)
Ancient bacteria species colonizing land with early land plants (Phys.org 11Sep2023)
Strullu-Derrien et al. (2023) Hapalosiphonacean cyanobacteria (Nostocales) thrived amid emerging embryophytes in an early Devonian (407-million-year-old) landscape