The Middle Devonian (393—383 Ma) occurs after the Emsian of the Early Devonian, and before the Frasnian of the Late Devonian.
The Middle Devonian is a fascinating time in plant evolution. During this time, plants begin to become more structurally complex, with elaborate anatomy, morphology, and increasing height. There are several groups and forms that appear on the landscape: ferns, "trees", woody vines, and possibly horsetails and proto-seed plants.
Above: the arrangement of continents during the Middle Devonian (~390 Ma)
Plants are becoming taller and more complex in anatomy, branching, and reproduction strategies
Spore-bearing plants radiate during this time
Some rhyniophytes, cooksonioids, and zosterophylls still exist in the landscape, but are quickly decreasing in dominance
Advanced clubmosses, protolepidodendrids, found in wet/swamp-like conditions
Early euphyllophytes, such as Chaleuria †, Douaphyton †, Kunia †, Oocampsa †, Pertica †, and Plantanophyton †, are found in the landscape
This includes the early fern-like plants, the rhacophytes, as well as iridopterids, which are possibly the ancestors of the horsetails or other equisetophytes
Pseudosporochnalean cladoxylopsids diversify
First tree-like plants appear in the Eifelian with Calamophyton † that stood 6-13 feet (2-4 meters) high
In the Givetian, Eospermatopteris † "trees" are over 25 feet (8m) tall.
Aneurophytalean progymnosperms appear, growing as woody vines among taller plants
Seed-megaspores or para-seeds appear during this time (e.g., Runcaria †, Granditetrasporites †, and Spermasporites †), which may be the ancestors to seed plants