The Tournaisian Age (359–347 Ma) is the first age in the Mississippian Epoch of the Carboniferous Period, occurring after the Famennian age (of the Devonian Period), and before the Visean Age.
358.9±0.4–346.7±0.4 Ma
Above: Sanfordiacaulis densifolia reconstruction (Gastaldo et al., 2024)
Below: Reconstruction of continent positions during Tournaisian
Several groups begin to disappear from the landscape: cladoxylopsids, Stenokoleales, and archaeopterids,
Sanfordiacaulis densifolia, a single-stemmed, non-woody "tree" estimated to be 20 feet tall (Gastaldo et al., 2024)
It possessed more than 250 leaves around its trunk, with partially-preserved leaves extending 1.75 meters, and probably another meter in real life
Affinities for this plant are unknown
Tropical Everwet and Summerwet
Dominated by arborescent lycopods, sphenophyllophytes, equisetophytes, and pteridosperms (Cleal and Thomas, 1991)
Clear increase in fossil charcoal in marine black shales during this time (Rimmer et al., 2015)
Burned plant material from ground fires was transported into watersheds
This is probably due to increased oxygen levels from the increased amount of plant life on land
Subtropical Desert
no fossil evidence
Warm Temperate
non-arborescent lycopsids and equisetophytes
Cool-cold temperate, arctic, and glacial
no fossil evidence
Earliest reptile footprints rewrite the timeline of tetrapod evolution (Phys.org 14May2025)
└Earliest amniote tracks recalibrate the timeline of tetrapod evolution (Long et al., 2025)
Rare tree fossils preserved with leaves unlike any plant known today (Live Science 2Feb2024)
└Enigmatic fossil plants with three-dimensional, arborescent-growth architecture from the earliest Carboniferous of New Brunswick, Canada (Gastaldo et al., 2024)