Evolutionary and Behavioural Ecology Group
in Sokendai, Japan

Welcome to Nobuyuki Kutsukake’s website

Lecturer, Department of Evolutionary Studies of Biosystems, The Graduate University for Advanced Studies

Hayama, Miura, Kanagawa, 240-0193, Japan

Tel: +81-46-858-1562, fax: +81-46-858-1544

email: kutsu (a) soken.ac.jp

Research Interests

I am a behavioral ecologist based in Tokyo, Japan.  I have been fascinated to animal lives since my childhood and am currently studying lovely mammals (primates, carnivores, and rodents), mainly in field.  Although my interests range wide areas of social evolution and social behaviour, I am particularly interested in understanding conflict resolution, communication and negotiation among group members in social living mammals, from both ultimate and proximate perspectives. 

Recently, I started a new project on general models of phenotypic evolution.  The progress is very slow but I hope to learn and find some interesting frameworks for contributing to our general understanding of phenotypic evolution. One output is a new phylogeny-based method to test an adaptive hypothesis.


Feel free to email me if you have any questions about my studies or if you are interested in collaboration.


Ongoing Projects

1. Socio-endocrinology in captive chimpanzees using saliva steroids (2004-, data collection finished)

2. Phenotypic evolution on phylogeny (2009-)

3. Social behaviour in fish (2010-)

4. Individual-based model for understanding animal sociality (2011-)

5. Supervision of graduate students' projects (too many, 2007-)


Completed Projects

1. Dominance and conflict resolution in wild Japanese macaques (1997-2000)

2. Social anxiety and conflict resolution in wild chimpanzees (2000-2001)

3. Greeting behaviour in captive black-and-white colobus (2003-2004)

4. Comparative analyses of reproductive skew in male primates (2004-2006)

5. Reproductive conflict and social behaviour in cooperatively breeding meerkats (2003-2010)

6. Group decision-making in eusocial naked mole-rats (2006-2012)


Selective Publications (you can download all publications from here)

Kutsukake N, Castles DL (2001) Reconciliation and variation in post-conflict stress in Japanese macaques (Macaca fuscata fuscata): testing the integrated hypothesis. Animal Cognition 4 :259-268

Kutsukake N, (2003) Assessing relationship quality and social anxiety among wild chimpanzees using self-directed behaviour. Behaviour 140: 1153-1171

Kutsukake N, Clutton-Brock TH (2006) Aggression and submission reflect reproductive conflict between females in cooperatively breeding meerkats. Behavioral Ecology and Sociobiology 59: 541-548

Kutsukake N (2006) The context and quality of social relationships affect vigilance behaviour in wild chimpanzees. Ethology 112: 581-591

Kutsukake N, Nunn CL (2006) Comparative tests of reproductive skew in male primates: the roles of demographic factors and incomplete control. Behavioral Ecology and Sociobiology 60: 695-706.

Kutsukake N, Clutton-Brock TH (2006) Social function of allogrooming in cooperatively breeding meerkats.Animal Behaviour 72: 1059-1068.

Kutsukake, N. & Clutton-Brock, T. H. (2008) The number of subordinates moderates intra-sexual competition among males in cooperatively breeding meerkats. Proceedings of the Royal Society of London. Series B, 

Biological Sciences 275: 209-216 

Kutsukake, N. & Clutton-Brock, T. H. (2008). Do meerkats engage in conflict management following aggression? Reconciliation, submission, and avoidance. Animal Behaviour. 75: 1441-1453

Kutsukake, N. (2009) Complexity, dynamics and diversity of sociality in group-living mammals. Ecological Research 24: 521-531.

Kutsukake, N. & Clutton-Brock TH (2010) Grooming and the value of social relationships in cooperatively breeding meerkats. Animal Behaviour 79: 271-279

Kutsukake N, Inada M, Sakamoto SH & Okanoya K. (2012) A distinct role of the queen in coordinated workload and soil distribution in eusocial naked mole-rats. PLOS ONE 7: e44584

Kutsukake N, Innan H. (2013) Simulation-based likelihood approach for evolutionary models of phenotypic traits on phylogeny. Evolution 67: 355-367