Ashkenazi Y-DNA Haplogroup E

Based upon the methodology posted here and using the sample set described here, as of January 2019 it appears that there are six ancestral Y-DNA lines in haplogroup E in the Ashkenazi Jewish population: (1) E-Y14891; (2) E-Y6923; (3) the E-BY932 subcluster of E-PF1975; (4) E-BY7450; (5) E-Z17697; and (6) E-BY11082.  Trees for each of those clusters are posted below.

E-Y14891
Upstream SNPs: 
M96>P147>M132>L499>P2>M215>M35>Z287>CTS10928>PF1962>M123>M34>L795> CTS4659>S21033>S11956>Y5427>Y14899>Y14891
ISOGG Tree (2019): E1b1b1b2a1a1a1ab2~
TMRCA (YFull Tree)
E-Y14891: 1,250 ybp (850-1,700 ybp @95%) 
E-Z36149: 650 ybp (350-1,000 ybp @95%) (E-Z36149 is at the same level as E-BY20366)
E-Y16781: 1,250 ybp (850-1,700 ybp @95%)
TMRCA (JewishDNA.net)
E1b-M84-Y14891 (AB-005): 460-708 CE (95%)

The E-Y14891 cluster follows the same pattern as many of the larger Ashkenazi clusters, with (1) a shared direct male ancestor dating back to about the second half of the first millenium CE, as evidenced by initial branching during that time frame; and (2) substantial branching within those subbranches that presumably dates back to the time when the Ashkenazi population began its massive expansion out of a small bottlenecked population, about 700 to 1,000 years ago.  

E-Y6923
Upstream SNPs: 
M96>P147>M132>L499>P2>M215>M35>Z287>CTS10928>PF1962>M34>Z841>Z849> CTS1727>L791>Y4971>Y6923
ISOGG Tree (2019): E1b1b1b2a1a4d2~
TMRCA (YFull Tree)
E-Y6923: 1,600 ybp (1,200-2,000 ybp @95%)
E-Y6938: 1,350 ybp (1,000-1,750 ybp @95%) (E-Y6923 is at the same level as E-Y6940)
E-B405: 1,250 ybp (1,000-1,550 ybp @95%)
E-A10736: 500 ybp (225-1,050 ybp @95%) (E-A10736 is at the same level as E-A10728)
E-Y15561: 1,000 ybp (700-1,300 ybp)
TMRCA (JewishDNA.net)
E1b-L91-Y6923 (AB-007): 673-861 CE (95%)

The E-Y6923 cluster follows the same pattern as many of the larger Ashkenazi clusters, with (1) a shared direct male ancestor dating back to about the second half of the first millenium CE, as evidenced by initial branching during that time frame; and (2) substantial branching within those subbranches that presumably dates back to the time when the Ashkenazi population began its massive expansion out of a small bottlenecked population, about 700 to 1,000 years ago.  

E-PF1975 (E-BY932 subcluster)
Upstream SNPs: 
M96>P147>M132>L499>P2>M215>Z827>CTS10298>PF1962>PF1975
ISOGG Tree (2019): E1b1b1b2a1b~
TMRCA (YFull Tree)
E-PF1975: 9,200 ybp (7,800-10,700 ybp @ 95% (identified as E-PF2025, on the same level as E-PF1975)
E-PF1974: 4,200 ybp (3,200-5,200 ybp @95%)
E-BY932: 1,450 ybp (900-2,000 ypb @ 95%) (identified as E-Y35934, on the same level as E-BY932)
TMRCA (JewishDNA.net)
E1b-Z830-PF1975 (AB-010): 721-1049 CE (95%)

The E-PF1974 branch of the tree above consists of a single branch -- FGC55987, with the upstream SNPs E-PF1952 and E-PF1974.  Branching is shown here starting at the E-PF1975 level to indicate that the E-FGC55987 branch and the E-BY932 branch share a direct male ancestor, who presumably lived in the Near East.

The E-BY932 cluster follows the same pattern as many of the larger Ashkenazi clusters, with (1) a shared direct male ancestor dating back to about the second half of the first millenium CE, as evidenced by initial branching during that time frame; and (2) substantial branching within those subbranches that presumably dates back to the time when the Ashkenazi population began its massive expansion out of a small bottlenecked population, about 700 to 1,000 years ago.  

E-BY7450

Upstream SNPs: 
M96>P147>M132>L499>P2>M215>M35>M78>Z1919>V22>Y2366>BY7446>BY7445> BY7447>BY44874>BY7452>BY7450
ISOGG Tree (2019): E1b1b1a1b2c1a1~
TMRCA (YFull Tree)
1,050 ybp (650-1,450 ybp @95%) (for E-BY7517, one level down from E-BY7450 and on the same level as E-BY7500)
TMRCA (JewishDNA.net)
E1b-V22-BY7500 (AB-001): 444-836 CE (95%) (for E-BY7500, one SNP downstream from BY7450)

The E-BY7450 cluster follows the same pattern as many of the larger Ashkenazi clusters, with (1) a shared direct male ancestor dating back to about the second half of the first millenium CE, as evidenced by initial branching during that time frame; and (2) substantial branching within those subbranches that presumably dates back to the time when the Ashkenazi population began its massive expansion out of a small bottlenecked population, about 700 to 1,000 years ago.  

E-Z17697

Upstream SNPs: 
E96>P147>M132>Z15511>M44>BY48020>Z17699>Z17697
TMRCA (YFull Tree)
1,000 ybp (550-1,500 ybp @95%)
TMRCA (JewishDNA.net)
?

The E-Z17697 cluster follows the same pattern as many of the larger Ashkenazi clusters, with (1) a shared direct male ancestor dating back to about the second half of the first millenium CE, as evidenced by initial branching during that time frame; and (2) substantial branching within those subbranches that presumably dates back to the time when the Ashkenazi population began its massive expansion out of a small bottlenecked population, about 700 to 1,000 years ago.  (Note that, for example, the E-Z17696 branch out of the E-Z17697 level leaves behind a E-Z17697* branch, and that, as a result, the tree above shows branching even though it shows only a single SNP downstream from E-Z17697.)

E-BY11082
Upstream SNPs: 
M96>P147>M132>L499>P2>M215>M35>Z827>L19>CTS10298>PF1962>M123>M34> L795>CTS4659>S12033>S11956>Y5427>PF6751>PF6748>S21355>PF6747>Z21014> BY1098>Z21018>BY11082
TMRCA (YFull Tree)
700 ybp (325-1,150 ybp) (for E-FGC32460, one SNP upstream from E-BY10750 and therefore presumably at the same level as E-BY11082)
TMRCA (JewishDNA.net)
?

Unlike the other Ashkenazi E Y-DNA clusters discussed above, based upon the current sample set the E-BY11082 cluster shows only a single ancestral branch prior to substantial branching within that branch (2) substantial branching within those subbranches that presumably dates back to the time when the Ashkenazi population began its massive expansion out of a small bottlenecked population, about 700 to 1,000 years ago.  

Y-DNA SNPs Not Appearing on Trees Posted Above
In addition to the SNPs posted above, there are a number of SNPs identified through the methodology used on this website that do not appear in any of the trees posted above. In some cases, primarily with regard to SNPs reported by Family Tree DNA based upon the results of Big Y testing or Geno 2.0 testing, the reported SNPs are ancestral to the SNPs set forth in one or more of the trees above; if the tested man is Ashkenazi on his direct male line, Big Y testing would likely determine that he belongs to one of the clusters set forth above. In other instances, it is possible that SNPs identify a small and/or undertested Ashkenazi cluster; further testing or information may identify such clusters.  Often, because the methodology used herein to identify potential Ashkenazi SNPs is overinclusive (especially for the 50 cM cohort but also, on occasion, for the 80 cM and 100 cM cohorts), SNPs set forth below do not reflect Ashkenazi (or Jewish) ancestry on the direct male line.  Finally, there are some SNPs (identified with "N/A") that do not appear on FTDNA's Y-DNA haplotree at all.

Upstream SNPs.  The following SNPs in the table above (identified in red) are upstream from SNPs found in one or more Ashkenazi Y-DNA clusters: L117 (not on the FTDNA Y-DNA Haplotree, but equivalent to M35), L791, L795, M123, M34, M35, M44, M78, M96, P2, PF6747, S11956, V22, and Z827.  

If men who are reported based upon STR or Geno 2.0 testing as having these SNPs are of Ashkenazi descent on their direct male line, there is a high probability that such men belong to one of those Ashkenazi Y-DNA clusters.  However, a large proportion of the men reported as having these terminal SNPs are not of Ashkenazi descent on their direct male lines.

Downstream SNPs that may reflect an Ashkenazi Y-DNA line.  The following terminal SNPs in the table above (identified in blue) do not appear in the Ashkenazi Y-DNA trees posted above, but may reflect an Ashkenazi Y-DNA line: BY11035, BY30352, BY5140, BY55895, BY77657, BY8528, MZ236, and S24513.  

Other SNPs.  With the exception of the SNPs that are not on FTDNA's Y-DNA Haplotree (identified with "N/A"), it is unlikely that men reported as having terminal SNPs other than the upstream SNPs identified above are of Ashkenazi descent on their direct male lines.