A regional assessment of white-tailed deer effects on plant invasion.  AoB Plants 10(1), plx047, doi:10.1093/aobpla/plx047

Averill, KM, Mortensen DA, Smithwick EAH, Kalisz S, McShea WJ, Bourg NA, Parker JD, Royo AA, Abrams MD, Apsley DK, Blossey B, Boucher CH, Caraher KL, DiTommaso A, Johnson SE, Masson R, Nuzzo VA


Interactive videos enhance learning about socio-ecological systems. Journal of Geography 117(1):40-49, 


BSmithwick EAH, Baxter E, Kim K, Edel-Malizia S, Rocco S, Blackstock D



Predicting root zone soil moisture with soil properties and satellite near-surface moisture data across the conterminous United States. Journal of Hydrology 546:393-404.

Baldwin D*, Manfreda S, Keller K, Smithwick EAH


A decade of colonization: the spread of the Asian Tiger Mosquito in Pennsylvania and Implications for disease risk. Journal of Vector Ecology 42(1): 3-12.
Taber ED*, Hutchinson, ML, Smithwick EAH, Blanford JI  



Grassland productivity in response to nutrient additions and herbivory is scale-dependent. PeerJ 4:e2745. 
Smithwick EAH, Baldwin DC*, Naithani KJ*


Build social costs into wildfire risks. Nature (Correspondence) 535 (7611): 231.
Smithwick, EAH

Generalist herbivore feeding selectivity is plant species-specific, regardless of nature/introduced status. Biological Invasions 18(5): 1247-1263. DOI: 10.1007/s10530-016-1063-z
Averill KM*, Mortensen DA, Smithwick EAH, Post E


Situated knowledge of pathogenic landscapes in Ghana: understanding the emergence of Buruli ulcer through qualitative analysis. Social Science and Medicine 150: 160-171. http://dx.doi.org/10.1016/j.socscimed.2015.12.005 

Tschakert P, Ricciardi V, Smithwick EAH, Machado M, Ferring D, Hausermann H, Bug L.



Influence of protected areas on malaria prevalence in sub-Saharan Africa. Applied Geography 64:35-45. DOI:10.1016/j.apgeog.2015.09.001

Taber E* and Smithwick EAH


Exploring interactions among multiple disturbance agents in forest landscapes: simulating effects of fire, beetles, and disease under climate change.  Ch.8, pp 201-231,  in Simulation Modeling of Forest Landscape Disturbances, Perera A., Remmel, R. and Buse L., eds., Springer International Publishing Switzerland, doi: 10.1007/978-3-319-19809-5_8.

Keane, RE, Loehman, R, Clark, J, Smithwick EAH, Miller C.


Buruli Ulcer disease and its association with land cover in southwestern Ghana.  PLoS Negl Trop Dis 9(6): e0003840.

Wu J, Tschakert P, Klutse E, Ferring D, Ricciardi V, Hausermann H, Oppong J, Smithwick EAH.

Representing Climate, Disturbance, and Vegetation Interactions in Landscape Models. Ecological Modelling  309-310: 33-47. 

Keane RE, McKenzie D, Falk DA, Smithwick EAH, Miller C, Kellog L-K B. 

Chemistry of natural waters and its relation to Buruli Ulcer in Ghana. Journal of Hydrology:  Regional Studies 3: 457-472.

Hagarty J,  Azanu D, Atosona , Voergbolo Smithwick EAH, Singha .

Sensitivity of four ecological models to adjustments in fine root turnover rate.  Ecological Modeling 297: 107-117.

McCormack, ML, Crisfield E, Raczka B, Schnekenburger F, Eissenstat DM, Smithwick EAH. 

Buruli Ulcer (BU) and landscape disturbance in Ghana 2007-2010.  Data set.  

Wu, J, Smithwick EAH, Ferring D, Hausermann H, Oppong J, Ricciardi V, Tschakert P. 

Available online [http://www.datacommons.psu.edu/ ] at The Pennsylvania State University, University Park, Pennsylvania, U.S.A.


Exploring interactions among multiple disturbance agents in forest landscapes: simulating effects of fire, beetles, and disease under climate change.  Ch.8 in Mapping and Modeling Forest Landscape Patterns, Perera A., Remmel, R. and Buse L., eds., Springer, in press.

Keane, RE, Loehman, R, Clark, J, Smithwick EAH, Miller C.


Improving the representation of roots in terrestrial models.  Ecological Modelling  291: 193–204

Smithwick EAHMcCormack MLSivandran G, Lucash MS.

Variability in root production, phenology, and turnover rate among 12 temperate tree species. Ecology, 95: 2224-2235

McCormack ML,  Adams TSSmithwick EAH, Eissenstat DM

Reconstructing disturbances and their biogeochemical consequences over multiple timescales. BioScience 64(2): 105-116

McLauchlan KK, PE Higuera, DG Gavin, SS Perakis, MC Mack, H Alexander, J Battles, F Biondi, B Buma, D Colombaroli, S Enders, DR Engstrom, FS Hu, JR Marlon, J Marshall, M McGlone, J L. Morris, LE Nave, BN Shuman, E Smithwick, DH Urrego, DA Wardle, CJ Williams, JJ Williams.  


Regional scale patterns of fine root lifespan and turnover under current and future climate.  Global Change Biology 19(6): 1697-1708. 

McCormack ML, Eissenstat DM, Prasad AM, Smithwick EAH

Root Stress and Nitrogen Deposition: Consequences and Research Priorities.  New Phytologist 197(3): 712-719.

Smithwick EAH, Eissenstat DM, Lovett G, Bowden R, Rustad L, and Driscoll C. 


Post-fire spatial patterns of soil nitrogen mineralization and microbial abundance.  PLoS ONE 7(11): e50597.

Smithwick EAH, Naithani KJ, Balser TC, Romme WH, and Turner MG. 

Contours of risk: spatializing human behaviour to understand disease dynamics in changing landscapes.  EcoHealth 9(3):251-5. 

Hausermann H, Tschakert P, Smithwick E, Ferring D, Amankwah R, Klutse E, Hagarty J, Kromel L.

Predicting fine root lifespan from functional traits in temperate trees.  New Phytologist Early View June 11.

McCormack ML, Adams TS, Smithwick EAH, Eissenstat DM.


Continued warming could transform Greater Yellowstone fire regimes by mid-21st Century. Proceedings of the National Academy of Science 108: 13165-13170.

Westerling AL, Turner MG, Smithwick EAH, Romme WH, Ryan MG. 

Variation in aboveground cover influences soil nitrogen availability at fine spatial scales following severe fire in subalpine conifer forests. Ecosystems 14(7):1081-1095.

Turner MG, Romme WH, Smithwick EAH, Tinker DB, Zhu J.

"Vulnerability of Landscape Carbon Fluxes to Future Climate and Fire in the Greater Yellowstone Ecosystem.” In: Questioning Greater Yellowstone’s Future: Climate, Land Use, and Invasive Species. Proceedings of the 10th Biennial Scientific Conference on the Greater Yellowstone Ecosystem. October 11–13, 2010, Mammoth Hot Springs Hotel, Yellowstone National Park. C. Andersen, ed., 131–134. Yellowstone National Park, WY, and Laramie, WY: Yellowstone Center for Resources and University of Wyoming William D. Ruckelshaus Institute of Environment and Natural Resources

Smithwick, E A H, Westerling A L, Turner MG, Romme WH, Ryan MG. 

Pyrogeography and biogeochemical resilience, pp. 143-164 in The Landscape Ecology of Fire, ed. by D. McKenzie, D. Falk, C. Miller, Springer.

Smithwick, EAH.

The wetland disturbance index: links with soil and water nitrate concentrations. Wetlands 31(5):853-863.

Cleveland MR, Smithwick EAH, Wardrop DH, Brooks RB. 


Pyrogeography:  Lessons for future northeastern U.S. landscapes. Penn State Environmental Law Review 18(2): 101-121.

Smithwick EAH.

Disturbance ecology, biogeochemistry and resilience: three decades of inquiry. Organized Oral Session 34. Bulletin of the Ecological Society of America 91:80-93.

Smithwick EAH.


Variation in foliar nitrogen and aboveground net primary production in young postfire lodgepole pine. Canadian Journal of Forest Research-Revue (Canadienne De Recherche Forestiere) 39:1024-1035.

Turner MG, Smithwick EAH, Tinker DB, and Romme WH. 

Long-Term Nitrogen Storage and Soil Nitrogen Availability in Post-Fire Lodgepole Pine Ecosystems. Ecosystems 12:792-806.

Smithwick EAH, Kashian DM, Ryan MG, and Turner MG. 

Modeling the effects of fire and climate change on carbon and nitrogen storage in lodgepole pine (Pinus contorta) stands. Global Change Biology 15:535-548.

Smithwick EAH, Ryan MG, Kashian DM, Romme WH, Tinker DB, and Turner MG.


Landscape heterogeneity following large fires: insights from Yellowstone National Park, USA. International Journal of Wildland Fire, in press.

Schoennagel T, Turner MG, and Smithwick EAH. 

Influence of coarse wood and pine saplings on nitrogen mineralization and microbial communities in young post-fire Pinus contorta. Forest Ecology and Management 256:59-67.

Metzger KL, Smithwick EAH, Tinker DB, Romme WH, Balser TC, and Turner MG.


Inorganic nitrogen availability following severe stand-replacing fire in the Greater Yellowstone Ecosystem. Inaugural paper, Proceedings of the National Academy of Science 104 (12): 4782-4789.

Turner, MG, Smithwick EAH, Metzger KL, Tinker DB, and Romme WH.


Editorial: Role of microbial communities in mediating ecosystem response to disturbance. Special Issue in Plant and Soil 289:1-3

Smithwick EAH.

Gentle introduction to complexity on landscapes.  Book Review for Ecology 87(11):2954-2955

Smithwick EAH.

Changing temporal patterns of forest carbon stores and net ecosystem carbon balance: The stand to landscape transformation, Landscape Ecology 22(1): 77-94. 

Smithwick EAH, Harmon ME, Domingo JB.


Spatial heterogeneity and soil nitrogen dynamics after severe fire in a black spruce (Picea mariana) forest, Alaska: Distinct controls at different scales. Biogeochemistry 76: 517-537 

Smithwick EAH, Mack MC, Turner MG, Chapin III FS, Zhu J, Balser TC. 

Post-fire soil nitrogen cycling in northern conifer landscapes affected by severe, stand-replacing fires.  Ecosystems 8: 163-181

Smithwick EAH, Turner MG, Mack MC, Chapin III, FS.

Variation in NH4+ mineralization and microbial communities with stand age in lodgepole pine (Pinus contorta) forests, Yellowstone National Park (USA). Soil Biology and Biochemistry 37: 1546-1559

Smithwick EAH, Turner MG, Metzger KL, Balser TC.

What the soil reveals: Maximum ecosystem C stores of the Pacific Northwest region, USA. Forest Ecology and Management 220: 270-283

Homann PS, Harmon ME, Remillard SM, Smithwick EAH.


Spatial Extrapolation: the science of predicting ecological patterns and processes. Bioscience 54(4): 310-320

Miller JM, Turner MG, Smithwick EAH, Dent LC, Stanley EH.


Modeling multi-scale effects of light limitations and edge-induced mortality on carbon stores in forest landscapes. Landscape Ecology 18(7): 701-721

Smithwick EAH, Harmon ME, Domingo JB.


Seasonal variation and ecosystem dependence of emission factors for selected trace gases and PM2.5 for southern African savanna fires. Journal of Geophysical Research 108(D24), 4758

Korontzi S, Ward DE, Susott RA, Yokelson RJ, Justice CO, Hobbs PV, Smithwick EAH, Hao WM. 


Potential upper bounds of carbon stores in the Pacific Northwest. Ecological Applications 12(5): 1303-1317

Smithwick EAH, Harmon ME, Acker SA, Remillard SM. 


Seasonality of carbon emissions from biomass burning in a Zambian savanna. Journal of Geophysical Research 104: 13,841 – 13,853.

Hoffa EA, Ward DE, Hao WM, Susott RA, Wakimoto RH.