Welcome to my website!

I am a biodiversity investigator with a background in terrestrial ecology.

My main goal is to elucidate how biodiversity patterns emerge and what mechanisms are involved in order to predict how environmental changes will affect diversity at different levels of organization.

I've been using spider assemblages as a biological model to understand the major drivers of biodiversity patterns at multiple spatial scales.

I am particularly interested in two biological systems: coastal dunes and oceanic island systems.

My research interests include:
  1. Biogeography and Macroecology
  2. Numerical ecology
  3. Dune ecology and conservation
  4. Island ecology and biogeography
I've been working as an associate researcher with the Centre of Molecular and Environmental Biology (University of Minho) and the Azorean Biodiversity Group (UNiversity of Azores)


21. Cardoso, P., Carvalho, J.C., Crespo, L.C., Arnedo, M.A. (2016) Optimal inventorying and monitoring of taxon, phylogenetic and functional diversity. bioRxiv; doi: http://dx.doi.org/10.1101/060400

20. Hintze, F., Duro, V., Carvalho, J.C., Eira, C., Rodrigues, P. & Vingada, J. (in press) Influence of reservoirs created by small dams on the activity of bats. Acta Chiropterologica.

19. Cardoso, P., Borges, P.A.V., Carvalho, J.C., Rigal, F., Gabriel, R., Cascalho, J. & Correia, L. (2015) Automated discovery of relationships, models and principles in ecology. bioRxiv; doi: http://dx.doi.org/10.1101/027839

18. Monteiro, S.S., Mendez-Fernandez, P., Piertney, S., Moffat, C.F., Ferreira, M., Vingada, J.V., Lopez, A., Brownlow, A., Jepson, P., Mikkelsen, B., Niemeyer, M., Carvalho, J.C., Pierce, G.J. (2015) Long-finned pilot whale population diversity and structure in Atlantic waters assessed through biogeochemical and genetic markers. Mar Ecol Prog Ser 536:243-257

17. Carvalho, J.C., Cardoso, P., Rigal, F., Triantis, K.A. & Borges, P.A.V. (2015) Modeling directional spatio‐temporal processes in island biogeography. Ecology and Evolution 5: 4671-4682

16. Cardoso, P., Rigal, F. & Carvalho, J.C. (2015) BAT–Biodiversity Assessment Tools, an R package for the measurement and estimation of alpha and beta taxon, phylogenetic and functional diversity. Methods in Ecology and Evolution, 6: 232-236.

15. Carvalho, J.C. & Cardoso, P. (2014) Drivers of beta diversity in Macaronesian spiders in relation to dispersal ability. Journal of Biogeography, 41: 1859-1870.

14. Cardoso P., Rigal, F., Borges, P.A.V. & Carvalho, J.C. (2014) A new frontier in biodiversity inventory: a proposal for estiamtors of phylogenetic and functional diversity. Methods in Ecology and Evolution, 5: 452-461.

13. Cardoso P., Rigal, F., Carvalho, J.C., Fortelius, M., Borges, P.A.V., Podani, J. & Schmera D. (2014) Partitioning taxon, phylogenetic and functional beta diversity into replacement and richness difference components. Journal of Biogeography, 41: 749-761

12. Boieiro, M.,  Carvalho, J.C., Cardoso, P.,  Aguiar, C.A.S., Rego, C., Silva, I.F., Amorim, I.R., Pereira, F., Azevedo, E.B., Borges, P.A.V. & Serrano, A.A.S. (2013) Spatial Factors Play a Major Role as Determinants of Endemic Ground Beetle Beta Diversity of Madeira Island Laurisilva.  PLoS ONE 05/2013; 8(5):e64591.

11. Carvalho, J.C., Cardoso, P., Borges, P.A.V., Schmera, D. & Podani, J. (2013) Measuring fractions of beta diversity and their relationships to nestedness: a theoretical and empirical comparison of novel approaches. Oikos, 122: 825–834.

10. Carvalho, J.C., Cardoso, P. & Gomes, P. (2012)  Determining the relative roles of species turnover and species richness differences in generating beta-diversity patterns. Global Ecology and Biogeography, 21: 760-771.

9. Carvalho, J.C., Cardoso, P., Crespo, L.C., Henriques, S., Carvalho, R. & Gomes, P. (2012) Determinants of spider species richness in coastal dunes along a gradient of mediterraneity. Insect Conservation and Diversity, 5: 127-137.

8. Carvalho, J.C., Cardoso, P., Crespo, L.C., Henriques, S., Carvalho, R. & Gomes, P. (2011) Determinants of beta diversity of spiders in coastal dunes along a gradient of mediterraneity. Diversity and Distributions, 17, 225-234.

7. Carvalho, J.C., Cardoso, P., Crespo, L.C., Henriques, S., Carvalho, R. & Gomes, P. (2011) Biogeographic patterns of spiders in coastal dunes along a gradient of mediterraneity. Biodiversity and Conservation, 20:873-894.

6. Pekár, S., Cardoso, P., Barriga, J. & Carvalho, J.C. (2011) Update to the zodariid spider fauna of the Iberian Peninsula and Madeira (Araneae: Zodariidae). Zootaxa, 2814: 19-32.

5. Torres, R.T., Carvalho, J.C., Panzacchi, M., Linnell, J.D.C. & Fonseca, C. (2011) Comparative habitat use of roe deer and moose in a human-modified landscape in southeastern Norway during winter. Ecological Research, 26:781-789

4. Carvalho, J.C. & Gomes P. (2004b). Feeding resources partitioning among four sympatric carnivores in the Peneda-Gerês National Park (Portugal). Journal of Zoology, 263: 275-283.

3. Carvalho, J.C. & Gomes P. (2004a). Influence of herbaceous cover, shelter and land cover structure on wild rabbit abundance in NW Portugal. Acta Theriologica, 49(1): 63-74.

2. Carvalho, J.C. & Gomes P. (2003). Habitat suitability for european wild rabbit (Oryctolagus cuniculus) with implications for restocking. Game and Wildlife Science, 20(4): 287-301.

1. Carvalho, J.C. & Gomes P. (2001). Food habits and trophic niche overlap of the red fox, European wild cat and common genet in the Peneda-Gerês National Park. Galemys, 13(2): 39-48.