Vermont Bryophyte List and Atlas               

Dorothy J. Allard                                         Last edited November 1, 2018

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The Vermont Bryophyte List has been developed over the last ten or so years from private and public collections and the literature. The list contains all bryophytes known from Vermont, although not all species known to the author only from herbarium specimens have been verified. Also included are synonyms (indented names), and global and state conservation ranks. For an explanation of ranks, see Natureserve Ranking Criteria.

Nomenclature follows Flora of North America, Vol. 27 (2007, Flora of North America Editorial Committee, Oxford University Press) and Vol. 28 (2014). For species to be published in Volume 29, nomenclature follows

The current list is available to view or download at the bottom of this page. First posted edition: December 27, 2007. 

Recent changes are described below; for older changes, click here.

The Vermont Bryophyte Atlas was developed from the same data sources: major herbaria containing Vermont collections (such as NY, VT, DUKE, NYS, FH, NEBC) and from numerous private collections or other data sources. The Atlas was originally created in version 3.2 of ArcView and is slowly being converted into ArcView 10 and released here as a collection of pdf files. In the current version, the town distribution of each Vermont species is displayed on a map showing extant (post-1960) and historic (pre-1960) records. Only the most recent data for the towns are displayed. An attribute table accompanies each map; it contains information on the data source, date of collection, collector, etc. Contents of the attribute tables are available upon request; please email To see the maps, click here.

October 31, 2018 changes: Campylostelium saxicola added to the list. State rank change for Jungermannia hyalina.

June 11, 2018 changes:  State rank changes for Bucklandiella sudetica, Grimmia donniana, and Splachnum pensylvanicum (all SH-->S1). Addition of Pseudoleskeella tectorum, Barbilophozia lycopodioides, Cephalozia ambigua, Cephaloziella elegans, and Cephaloziella rubella var. bifida; Cephaloziella arctica removed. A few liverwort names were updated. Splachnum pennsylvanicum corrected to Splachnum pensylvanicum.

February 12, 2018 changes: Lophozia silvicola, Lophozia ventricosa var. longiflora, and Lophozia ventricosa var. ventricosa added; Lophozia ventricosa and Tortella tortuosa var. fragilifolia removed.

February 4, 2018 changes: State rank changes for Tortella tortuosa var. fragilifolia and Porella pinnata.

December 16, 2017 changes: Tortella tortuosa var. fragilifolia added back to the list (note that it was removed back in 2010); state rank changes for Abietinella abietina, Amphidium mougeotii, Dicranella schreberiana, Lewinskya sordida, and Orthotrichum stellatum.

November 7, 2017 changes: Rank changes for Trematodon longicollis and Pohlia lescuriana.

February 5, 2017 changes: Rank changes for Lewinskya sordida, Fissidens bryoides and Dicranum spurium.

December 31, 2016 changes: Rank changes for Anacamptodon splachnoides and Andreaea rothii. Three species of Orthotrichum moved to Lewinskya. Brachythecium calcareum synonymized with B. campestre.

November 22, 2016 changes: Two new taxa, Ephemerum crassinervium ssp. crassinervium and Physcomitrium collenchymatum added. Rank for Lophozia wenzelii changed from SH to S1. Ulota crispa removed; Ulota crispula added.

October 13, 2016 changes: Two new taxa, Scapania carinthiaca and Tetraplodon angustatus added. Rank change for Tetraplodon mnioides. Riccia cavernosa (the currently accepted name) replaces R. crystallina, and the rank is updated.

Nov. 14, 2015 changes: State rank changes for numerous species. Hornworts separated from liverworts and included at the end of the list. Two new taxa, Bracythecium falcatum and Zygodon conoideus added.

April 3, 2011 changes: Taxon removed from list: Archidium alternifolium (Hedw.) Schimp. (it was represented by a single mis-identified specimen). State rank changes for Acaulon muticum var. rufescens, Atrichum crispulum, Barbula convoluta var. convoluta, Didymodon fallax, and Homomallium adnatum. Global ranks reviewed and adjusted to match those in NatureServe Explorer. State ranks corrected to eliminate T-ranks and Q-ranks. Three typographical errors in species names corrected.

January 12, 2011 changes: Taxon removed from list: Atrichum undulatum (James) Sull. Taxon added to list: Pohlia sphagnicola. State rank changes for Bryoandersonia illecebra, Helodium paludosum, Limprichtia cossonii, Sphagnum isoviitae, Calypogeia sphagnicola, Cephalozia elachista, Cladopodiella fluitans, Riccardia palmata, and Scapania paludicola.

November 5, 2010 changes: Taxa removed from list: Tortella tortuosa var. fragilifolia, Tortella tortuosa var. tortuosa. Taxa added to list: Tortella bambergeri, Drepanocladus longifolius. State rank changes for Hypnum cupressiforme var. filiforme, H. cupressiforme s.l., H. cupressiforme var. cupressiforme, H. cupressiforme var. resupinatum, Brachythecium populeum, Mnium spinulosum, Fissidens taxifolius, Taxiphyllum deplanatum, Codriophorus acicularis, Schistidium liliputanum, Ditrichum pallidum, Pohlia elongata var. elongata, Lophozia ventricosa s.l., Lophozia ventricosa var. silvicola, Lophozia ventricosa var. longiflora.

Campylostelium saxicola, Sterling Falls Gorge, Stowe, Vermont
Camplyostelium saxicola, Sterling Falls Gorge, Stowe, Vermont

Dorothy Allard,
Nov 1, 2018, 7:39 AM