Régis Chirat

    Université Lyon 1
    27–43 Boulevard du 1
1 Novembre 1918,
    F-69622 Villeurbanne Cedex, France.

                                                                                                                                   Knemiceras uhligi

Research interests
  • Mechanical basis of seashells morphogenesis and evolution ("Devo-Evo")
  • Nautiloid paleobiology 

  • Rudraraju S., Moulton D. E., Chirat R., Goriely A., Garikipati K. (Submitted) A computational framework for the morpho-elastic development of molluskan shells by surface and volume growth [available on arXiv]
  • Erlich E., Howell R., Goriely A., Chirat R., Moulton D. E. (2018).- Mechanical feedback in seashell growth and form. The ANZIAM Journal. 59: 581-606. special issue on ‘Mechanics in Biology’. pdf
  • Moulton D. E., Goriely A. & Chirat R. (2018).- Comment les coquillages acquièrent leur forme. Pour la Science, 491: 50-57. pdf
  • Moulton D. E., Goriely A. & Chirat R. (2018).- How seashells take shape. Scientific American, 318, 4: 68-75. pdf. Highlighted here and here
  • Erlich A., Moulton D. E., Goriely A. & Chirat R. (2016).- Morphomechanics and developmental constraints in the evolution of ammonites shell form. Journal of Experimental Zoology Part B: Molecular and Developmental Evolution, 326: 437-450. pdf. See the interactive model here
  • Moulton D. E., Goriely A. & Chirat R. (2015).- The morpho-mechanical basis of ammonite form. Journal of Theoretical Biology, 364: 220-230. pdf. Highlighted here and here.
  • Chirat R., Moulton D. E. & Goriely A. (2013).- Mechanical basis of morphogenesis and convergent evolution of spiny seashells. Proceedings of the National Academy of Sciences of the USA, 110 (15): 6015-6020. pdf. Highlighted in Nature and The Scientist
  • Moulton D. E., Goriely A. & Chirat R. (2012).- Mechanical growth and morphogenesis of seashells. Journal of Theoretical Biology, 311: 69-79. pdf
  • Schlogl J., Chirat R.,  Balter V., Joachimski M., Hudácková N. & Quillévéré F. (2011).- Aturia from the Miocene Paratethys: an exceptional window on nautilid habitat and lifestyle.Palaeogeography, Palaeoclimatology, Palaeoecology 308, 330-338. pdf
  • Urdy S., Goudemand N., Bucher H. & Chirat R. (2010).- Growth-dependent phenotypic variation of molluscan shells: implications for allometric data interpretation. Journal of Experimental Zoology Part B: Molecular and Developmental Evolution, 314: 303–326. pdf
  • Urdy S., Goudemand N., Bucher H. & Chirat R. (2010).- Allometries and the morphogenesis of the molluscan shell: a quantitative and theoretical model. Journal of Experimental Zoology Part B: Molecular and Developmental Evolution, 314: 280–302. pdf
  • Chirat R., Enay R., Hantzpergue P. & Mangold C. (2008).- Developmental integration related to buoyancy control in nautiloids. Evidence from unusual septal approximation and ontogenetic allometries in a jurassic species. Palaeontology, 51, 1: 251–261. pdf
  • Urdy S. & Chirat R. (2006).- Snail shell coiling (re-)evolution and the evo-devo revolution. Journal of Zoological Systematics and Evolutionary Research.44(1), 1–7. pdf
  • Chirat R. & Bucher H. (2006)- Shell microstructure and morphogenesis of the ornamentation in Cymatoceras Hyatt, 1883, Cretaceous Nautilida. Systematic implications.  Lethaia, 39: 57-64. pdf
  • Chirat R., Vaslet D. & Le Nindre Y.-M. (2006).- Nautiloids from the Permian-Triassic Khuff Formation, central Saudi Arabia. GeoArabia, v. 11, no. 1: 81-92.pdf
  • Bucher H., Chirat R. & Guex J. (2003).- Morphogenetic origin of radial lirae and mode of shell growth in Calliphylloceras (Jurassic Ammonoidea). Eclogae Geologicae Helvetiae. 96: 495-502. pdf
  • Chirat R. & Boletzky S. von (2003).- Morphogenetic significance of the conchal furrow in nautiloids : evidence from early embryonic shell development of Jurassic Nautilida. Lethaia, 36 : 161-170. pdf
  • Chirat R. (2001).- Anomalies of embryonic shell growth in post-Triassic Nautilida. Paleobiology 27 : 485-499. pdf
  • Chirat R. (2000).- The so-called “ cosmopolitan palaeobiogeographic distribution ” of Tertiary Nautilida of the genus Aturia Bronn 1838 : the result of post-mortem transport by oceanic palaeocurrents. Palaeogeography, Palaeoclimatology,Palaeoecology, 157 : 59-77. pdf
  • Rioult M. & Chirat R. (1999 ).- Ammonites jurassiques sur la bordure occidentale du Bassin anglo-parisien. Contrôle et modalités de leur répartition. Bull. Soc. Linn. Normandie, 117 : 49-60 . 
  • Chirat R. & Rioult M. (1998).- Occurrence of early post-hatching Jurassic Nautilida in Normandy, France. Palaeobiologic, palaeoecologic and palaeobiogeographic implications. Lethaia, 31 : 137-148. pdf
  • Chirat R. & Rioult M. (1998).- Les reliefs de bordures de massifs anciens : des sites privilégiés de reproduction pour les Nautilida Jurassiques. Comptes Rendus de l'Académie des Sciences, 327 : 197-202. pdf
  • Chirat R. (1997).- Les Nautilida post-triasiques. Paléobiologie, paléoécologie, paléobiogéographie et systématique à la lumière des données récemment acquises sur Nautilus. PhD Thesis, University of Caen. 322 pp.