Reprints Also Available Upon Request
Journal Articles

Love, A.C., T.A. Stewart, G.P. Wagner, and S.A. Newman. 2017. Perspectives on integrating genetic and physical explanations of evolution and development. Integrative and Comparative Biology. doi: 10.1093/icb/icx121.

Love, A.C. and M.J. Nathan. 2015. The idealization of causation in mechanistic explanation. Philosophy of Science 82:761–774 . [journal pdf] [preprint pdf]

Love, A.C. 2015. Collaborative explanation, explanatory roles, and scientific explaining in practice. Studies in History and Philosophy of Science 52:88–94. [journal pdf] [preprint pdf]

Moczek, A.P., K.E. Sears, A. Stollewerk, P.J. Wittkopp, P. Diggle, I. Dworkin, C. Ledon-Rettig, D.Q. Mattus, D.Q., S. Roth, E. Abouheif, F.D. Brown, C-H. Chiu, C.S. Cohen, A.W. De Tomaso, S.F. Gilbert, B.K. Hall, A.C. Love, D.C. Lyons, T. Sanger, J. Smith, C. Specht, M. Vallejo-Marin, and C.G. Extavour. 2015. The significance and scope of evolutionary developmental biology: a vision for the 21st century. Evolution & Development 17:198–219. [journal pdf]

O’Malley, M.A., I. Brigandt, A.C. Love, J.W. Crawford, J.A. Gilbert, R. Knight, S.D. Mitchell, and F. Rohwer. 2014. Multilevel research strategies and biological systems. Philosophy of Science. 81:811–828. [journal pdf] [preprint pdf]

Love, A.C. and G.L. Lugar 2013. Dimensions of integration in interdisciplinary explanations of the origin of evolutionary novelty. Studies in the History and Philosophy of Biological and Biomedical Sciences. 44:537550. [journal pdf] [preprint pdf]

Love, A.C. 2013. Theory is as theory does: scientific practice and theory structure in biology”, Biological Theory 7:325337. [journal pdf] [Spring SharedIT link] [preprint pdf] [erratum]

Love, A.C. and M. Travisano 2013. Microbes modeling ontogeny. Biology & Philosophy 28: 161188. [journal pdf] [Springer SharedIT Link] [preprint pdf]

Love, A.C. 2013. Interdisciplinary lessons for the teaching of biology from the practice of Evo-devo. Science & Education 22:255278. [journal pdf] [Springer SharedIT Link] [preprint pdf]

Brigandt, I. and A.C. Love 2012. Conceptualizing evolutionary novelty: moving beyond definitional debates. J. Exp. Zool. (Mol. Dev. Evol.) 318B:417–427. [journal pdf] [preprint pdf]

Love, A.C. 2012. Hierarchy, causation and explanation: ubiquity, locality, and pluralism. Interface Focus 2:115–125. [journal pdf] [preprint pdf]

Hüttemann, A. and A.C. Love 2011. Aspects of reductive explanation in biological science: intrinsicality, fundamentality, and temporality. British Journal for Philosophy of Science 62:519–549. [journal pdf] [preprint pdf]

Brigandt, I. and A.C. Love 2010. Evolutionary novelty and the Evo-devo synthesis: field notes. Evolutionary Biology 37:93–99. [journal pdf] [Springer SharedIT Link] [preprint pdf]

Love, A.C. 2010. Darwin’s ‘imaginary illustrations’: creatively teaching evolutionary concepts and the nature of science. The American Biology Teacher 72:82–89. [journal pdf] [preprint pdf]

Love, A.C. 2010. Idealization in evolutionary developmental investigation: a tension between phenotypic plasticity and normal stages. Philosophical Transactions of the Royal Society B 365:679–690. [journal pdf] [preprint pdf]

Love, A.C. 2009. Marine invertebrates, model organisms, and the modern synthesis: epistemic values, evo-devo, and exclusion. Theory in Biosciences 128:19–42. [journal pdf] [Springer SharedIT Link] [preprint pdf]

Love, A.C. 2009. Typology reconfigured: from the metaphysics of essentialism to the epistemology of representation. Acta Biotheoretica 57:51–75. [journal pdf] [preprint pdf]

Love, A.C. 2008. Explaining evolutionary innovation and novelty: criteria of adequacy and multidisciplinary prerequisites. Philosophy of Science 75:874–886. [journal pdf] [preprint pdf]

Love, A.C., A.E. Lee, M.E. Andrews, and R.A. Raff 2008. Co-option and dissociation in larval origins and evolution: the sea urchin larval gut. Evolution & Development 10:74–88. [journal pdf] [preprint pdf]

Love, A.C. 2008. From philosophy to science (to natural philosophy): evolutionary developmental perspectives. The Quarterly Review of Biology 83:65–76. [journal pdf] [preprint pdf]

Love, A.C. 2007. Functional homology and homology of function: biological concepts and philosophical consequences. Biology & Philosophy 22:691–708. [journal pdf] [Springer SharedIT Link] [preprint pdf]

Love, A.C., M.E. Andrews, R.A. Raff 2007. Gene expression patterns in a novel animal appendage: the sea urchin pluteus arm. Evolution & Development 9:51–68. [journal pdf] [preprint pdf]

Love, A.C. 2006. Evolutionary morphology and evo-devo: hierarchy and novelty. Theory in Biosciences 124:317–333. [journal pdf] [preprint pdf]

Love, A.C. 2006. Reflections on the middle stages of evo-devo's ontogeny. Biological Theory 1:94–97. [journal pdf]

Love, A.C. and R.A. Raff 2006. Larval ectoderm, organizational homology, and the origins of evolutionary novelty. Journal of Experimental Zoology (Mol Dev Evol) 306B:18–34. [journal pdf] [preprint pdf]

Ingram, E.L., E. Lehman, A.C. Love, and K.M. Polacek 2004. Fostering inquiry in nonlaboratory settings. Journal of College Science Teaching 34(1):39–43. [journal pdf]

Raff, R.A. and A.C. Love 2004. Kowalevsky, comparative evolutionary embryology, and the intellectual lineage of evo-devo. Journal of Experimental Zoology (Mol Dev Evol) 302B:19–34. [journal pdf] [preprint pdf]

Love, A.C. 2003. Evolvability, dispositions, and intrinsicality. Philosophy of Science 70:1015–1027. [journal pdf] [preprint pdf]

Love, A.C. and R.A. Raff 2003. Knowing your ancestors: themes in the history of evo-devo. Evolution & Development 5:327–330. [journal pdf] [preprint pdf]

Love, A.C. 2003. Evolutionary morphology, innovation, and the synthesis of evolutionary and developmental biology. Biology & Philosophy 18:309–345. [journal pdf] [Springer Shared It Link] [preprint pdf]

Translated and reprinted

  • Love, A.C. 2013 [2003]. Morfologia evoluzionistica, innovazione e sintesi della biologia evoluzionistica e dello sviluppo. In A. Pinotti and S. Tedesco (eds.). Estetica e science della vita: Morfologia, biologia teoretica, evo-devo. Milan: Raffaello Cortina Editore, 289–323.

Love, A.C. 2002. Darwin and Cirripedia prior to 1846: exploring the origins of the barnacle research. Journal of the History of Biology 35:251–289. [journal pdf] [Springer SharedIt Link] [preprint pdf]


Love, A.C. under contract. Evolution and Development. Cambridge University Press.

Love, A.C. (ed.) 2015. Conceptual Change in Biology: Scientific and Philosophical Perspectives on Evolution and Development. Boston Studies in Philosophy of Science (Volume 307). Berlin: Springer. [online/e-book]

Book Chapters and Related

Sarkar, S., A.C. Love, and W.C. Wimsatt. forthcoming. Reductionism in biology. Oxford Bibliographies in Philosophy.

Love, A.C. forthcoming. Combining genetic and physical causation in developmental explanations. In C.K. Waters and J. Woodward (eds), Causal Reasoning in Biology. Minnesota Studies in the Philosophy of Science. Minneapolis: University of Minnesota Press.

Love, A.C. forthcoming. Situating evolutionary developmental biology in evolutionary theory. In S.M. Scheiner and D.P Mindell (eds), The Theory of Evolution. University of Chicago Press.

Love, A.C. and R.R. Strathmann. forthcoming. Marine invertebrate larvae: model life histories for development, ecology, and evolution. In T.J. Carrier, A.M. Reitzel, and A. Heyland (eds), Evolutionary Ecology of Marine Invertebrate Larvae. Oxford University Press.

Love, A.C. forthcoming. Individuation, individuality, and experimental practice in developmental biology. In O. Bueno, R.-L. Chen, and M.B. Fagan (eds), Individuation across Experimental and Theoretical Sciences. Oxford University Press.

Love, A.C. 2018. Developmental mechanisms. In S. Glennan and P. Illari (eds), The Routledge Handbook of the Philosophy of Mechanisms. New York: Routledge, pp. 159–187.

Love, A.C. 2017. Evo-devo and the structure(s) of evolutionary theory: a different kind of challenge. In P. Huneman and D. Walsh (eds), Challenging the Modern Synthesis. New York: Oxford University Press, pp. 159–187.

Love, A.C. and I. Brigandt. 2017. Philosophical dimensions of individuality. In S. Lidgard and L.K. Nyhart (eds), Biological Individuality: Integrating Scientific, Philosophical, and Historical Perspectives. University of Chicago Press, pp. 318–348. [open access]

Love, A.C. 2017. Building integrated explanatory models of complex biological phenomena: From Mill’s methods to a causal mosaic. In M. Massimi and J.-W. Romeijn (eds), EPSA Philosophy of Science: Düsseldorf 2015. The European Philosophy of Science Association Proceedings, Vol. 5. Dordrecht: Springer, pp. 221–232.

Liu, K.E., A.C. Love, and M. Travisano. 2017. How cancer spreads: reconceptualizing a disease. In G. Boniolo and M.J. Nathan (eds), Philosophy of Molecular Medicine: Foundational Issues in Research and Practice. Routledge. pp. 100–121.

Love, A.C. 2016. Explaining the origins of multicellularity: between evolutionary dynamics and developmental mechanisms. In: K.J. Niklas and S.A. Newman (eds.), Multicellularity: Origins and Evolution. Cambridge, MA: MIT Press, 279–295.

Hüttemann, A. and A.C. Love. 2016. Reduction. In P. Humphries (ed), The Oxford Handbook of Philosophy of Science. New York: Oxford University Press, 460–484.

Love, A.C. and D. Urban. 2016. Developmental evolution of novel structures – animals. In R. Kliman (ed.), Encyclopedia of Evolutionary Biology. Volume 3. Oxford: Academic Press, 136–145. [published version]

Love, A.C. 2015. Developmental biology. The Stanford Encyclopedia of Philosophy.

Love, A.C. 2015. Conceptual change and evolutionary developmental biology. In A.C. Love (ed), Conceptual Change in Biology: Scientific and Philosophical Perspectives on Evolution and Development. Boston Studies in Philosophy of Science. Berlin: Springer, 1–54. [published version] [

Love, A.C. 2015. Evolutionary developmental biology: philosophical issues. In T. Heams, P. Huneman, L. Lecointre, and M. Silberstein (eds), Handbook of Evolutionary Thinking in the Sciences. Berlin: Springer. 265–283. [published version] [preprint pdf]

Love, A.C. 2014. The erotetic organization of developmental biology. In A. Minelli and T. Pradeu (eds), Towards a Theory of Development. Oxford: Oxford University Press. 33–55. [preprint pdf] formatted pdf available upon request

Love, A.C. 2013. Teaching evolutionary developmental biology: concepts, problems, and controversy. In K. Kampourakis (ed), The Philosophy of Biology: A Companion for Educators. Berlin: Springer, 323–341. [published version] [preprint pdf]

Love, A.C. 2012. Material versus formal theories in philosophy of science: a methodological interpretation. In H. de Regt, S. Okasha, and S. Hartmann (eds), EPSA Philosophy of Science: Amsterdam 2009. The European Philosophy of Science Association Proceedings, Vol. 1. Berlin: Springer, 175–185. [published version] [preprint pdf]

Love, A.C. and A. Hüttemann 2011. Comparing part-whole explanations in biology and physics. In D. Dieks, W.J. Gonzalez, S. Hartmann, T. Uebel, and M. Weber (eds), Explanation, Prediction, and Confirmation. Berlin: Springer, 183–202. [published version] [preprint pdf]

Love, A.C. 2011. Darwin’s functional reasoning and homology. In M. Wheeler (ed), 150 Years of Evolution: Darwin’s Impact on Contemporary Thought & Culture. San Diego: SDSU Press, 49–67. [preprint pdf] formatted pdf available upon request

Love, A.C. 2010. Rethinking the structure of evolutionary theory for an extended synthesis. In M. Pigliucci and G. Müller (eds) Evolution—The Extended Synthesis. Cambridge, MA: MIT Press, 403–441. [preprint pdf] formatted pdf available upon request

Brigandt, I. and A.C. Love 2008. Reductionism in biology. Stanford Encyclopedia of Philosophy. [substantive revision and update: 2012]

Love, A.C. 2008. Explaining the ontogeny of form: philosophical issues. In S. Sarkar and A. Plutynski (eds.) A Companion to the Philosophy of Biology. Malden, MA: Blackwell Publishers, 223–247. [preprint pdfformatted pdf available upon request

Love, A.C. 2007. Morphological and paleontological perspectives for a history of evo-devo. In M. Laubichler and J. Maienschein (eds.) From Embryology to Evo-Devo: A History of Developmental Evolution. Cambridge, MA: MIT Press, 267–307. [preprint pdf] formatted pdf available upon request

Love, A.C. 2002. Comments on Robert Brandon’s “The difference between drift and selection: A reply to Millstein” (web publication).