Shannon,G.,Angeloni, L.M., WITTEMYER, G., Fristrup, K.M., and Crooks, K.R. 2014. Roadtraffic noise modifies behaviour of a keystone species. Animal Behaviour 94:135-141

Northrup, J. M., Anderson, C. R., WITTEMYER, G. 2014. Effects ofcapture and handling on movement behavior of mule deer. Journal of Wildlife Management 78:731-738

Wall, J., WITTEMYER, G., Klinkenberg, B., and Douglas-Hamilton, I. 2014. Novel opportunities for wildlifeconservation and management using real-time monitoring. Ecological Applications 24:593-601

Chiyo, P.I., Grieneisen, L.E.,WITTEMYER, G., Moss, C.J., Lee, P.C., Douglas-Hamilton, I., Archie, E.A. 2014.The influence of social structure, habitat, and host traits on the transmissionof Escherichia coli in wildelephants. PLoS One e93408

Noa Pinter-Wollman, N., Hobson, E.A., Smith, J. E., Edelman, A. J., Shizuka, D., Waters, J. S., de Silva, S., Prager, S. D., Sasaki,T., WITTEMYER, G., Fewell, J., McDonald, D. B. 2014. The dynamics of network dynamics. BehavioralEcology 25:260-261

Noa Pinter-Wollman, N., Hobson, E.A., Smith, J. E., Edelman, A. J., Shizuka, D., Waters, J. S., de Silva, S., Prager, S. D., Sasaki,T., WITTEMYER, G., Fewell, J., McDonald, D. B. 2014. The dynamics of animal social networks: analytical, conceptual, andtheoretical advances. Behavioral Ecology  25:242-255

Sienne, J.M., Buchwald, R., WITTEMYER, G. 2014. Differentiation in mineral constituents in water and soil resources across Central African bais (forest clearings). European Journal of Wildlife Research 60:377-382

Gessner, J., Buchwald,R., WITTEMYER, G. 2014. Assessing species occurrence and species specific usepatterns of bais in Central Africa with camera traps. African Journal ofEcology 52:59-68.


Turkalo,A.K., Wrege, P., and WITTEMYER, G. 2013. Longterm monitoring of the Dzanga bai forest elephants: Population description and bai use patterns. PLoS One 8(12):e85154

deSilva, S.,Webber C.E., Weerathunga, U.S., Pushpakumara, T.V., Tharanga, J.S., Weerakoon,D., WITTEMYER, G. 2013. Demographicvariables for wild Asian elephants using longitudinal observations. PLoS One 8(12):e82788

PolanskyL., Douglas-Hamilton, I., WITTEMYER, G. 2013. Using diel movement behavior toinfer foraging strategies related to ecological and social factors in elephants. MovementEcology 1:13

Creel, S., Becker, M. S., Durant,S. M., M'Soka, J., Matandiko, W., Dickman, A. J., Christianson, D., Droge, E.,Mweetwa, T., Pettorelli, N., Rosenblatt, E., Schuette, P., Woodroffe, R.,Bashir, S., Beudels-Jamar, R. C., Blake, S., Borner, M., Breitenmoser, C.,Broekhuis, F., Cozzi, G., Davenport, T. R. B., Deutsch, J., Dollar, L.,Dolrenry, S., Douglas-Hamilton, I., Fitzherbert, E., Foley, C., Hazzah, L.,Henschel, P., Hilborn, R., Hopcraft, J. G. C., Ikanda, D., Jacobson, A.,Joubert, B., Joubert, D., Kelly, M. S., Lichtenfeld, L., Mace, G. M., Milanzi,J., Mitchell, N., Msuha, M., Muir, R., Nyahongo, J., Pimm, S., Purchase, G.,Schenck, C., Sillero-Zubiri, C., Sinclair, A. R. E., Songorwa, A. N.,Stanley-Price, M., Tehou, C. A., Trout, C., Wall, J., WITTEMYER, G.,Zimmermann, A. 2013. Conserving large populations of lions – the argument forfences has holes. Ecology Letters 16:1413-e3

Uno, K., Quade, J. Fisher, D.C., WITTEMYER, G., Douglas-Hamilton,I., Adanje, S., Omondi, P., Litoroh, M., Cerling, T.E. 2013. Bomb-curve radiocarbonmeasurements of recent biologic tissues and applications to stable isotopeecology and paleoecology, and to wildlife forensics. Proceedings of the National Academy of Sciences 110(29):11736-11741

Lynch, E., Angeloni, L., Fristrup, K.,Joyce, D., WITTEMYER, G.  2013. Theuse of on-animal acoustical recording devices for studying animal behaviour. Ecology and Evolution 3(7):2030-2037

Northrup,J. M., Hooten, M. B., Anderson, C. R., WITTEMYER, G. 2013. Practicalguidance on availability sampling for resource selection functions. Ecology 94(7):1456-1463

Maisels,F., Strindberg, S., Blake, S., WITTEMYER, G., Hart, J., Williamson, E., Aba'a,R., Abitsi, G., Ambahe, R., Amsini, F., Bakabana, P., Hicks, C., Bayogo, R.,Bechem, M., Beyers, R., Bezangoye, A., Bonyenge, A., Boudjan, P., Bout, N.,Akou, M.E., Bene Bene, L., Fosso, B., Greengrass,E.,  Grossmann, F., Ikamba-Nkulu, C., Ilambu, O.,Inogwabini, B., Iyenguet, F., Kiminou, F., Kokangoye, M.,Kujirakwinja, D., Latour, S., Liengola, I., Mackaya, Q., Madidi, J., Madzoke,B., Makoumbou, C., Malanda, G., Malonga, R., Mbani, O., Mbendzo, V., Mbenkira,M., Ambassa, E.,  Ekinde, A.,Mboh, H., Mihindou, Y., Morgan, B., Motsaba, P., Moukala, G., Mounguengui, A.,Mowawa, B., Ndzai, C., Nixon, S., Nkumu, P., Nzolani, F., Plumptre, A., Rainey,H., de Semoboli, B., Serckx, A., Stokes, E., Turkalo, A., Vanleeuwe, H., Vosper, A., Warren, Y., 2013.Devastating decline of forest elephants in Central Africa.  PLoS One Devastating declineof forest elephants in Central Africa.  PLoS One 8(3): e59469

Wall,J., WITTEMYER, G., Klinkenberg, B. LeMay, V., and Douglas-Hamilton, I. 2013. Characterizing properties and drivers of long distancemovements by elephants (Loxodontaafricana) in the Gourma, Mali. BiologicalConservation 157:60-68

Northrup,J.M. and WITTEMYER, G. 2013. Characterizingthe impacts of emerging energy development on wildlife, with an eye towardsmitigation. Ecology Letters 16:112-125

WITTEMYER,G., Daballen, D., and Douglas-Hamilton, I. 2013.Comparative demography of an at risk African elephant population. PLoS One 8(1):e53726


de Silva, S. and Wittemyer, G. 2012. Acomparison of social organization in Asian elephants and African savannahelephants. International Journal ofPrimatology 33:1125-1141.


Wittemyer, G., Daballen,D., and Douglas-Hamilton, I. 2011. Rising ivory prices threaten elephants. Nature 476:282-283.


Polansky, L., Wittemyer, G., Cross, P.C., Tambling, C.J. & Getz, W.M. 2010. Fourier and wavelet analyses applied to animal movement time series. Ecology 91:1506-1518.

Wasser, S., et al. 2010. Elephants, Ivory, and Trade. Science 327:1331-1332

Kahindi, O., Wittemyer, G., King, J., Ihwagi, F., Omondi, P., & Douglas-Hamilton, I. 2010. Employing participatory surveys to monitor the illegal killing of elephants across diverse land uses in Laikipia-Samburu, Kenya. African Journal of Ecology 48:972-983


Wittemyer, G., Cerling, T.E., & Douglas-Hamilton I. 2009. Establishing longitudinal diet chronologies from isotopic profiles in serially collected hairs. Chemical Geology 267:3-11 PDF

Wittemyer, G., Okello, J.B.A., Rasmussen, H.B., Arctander, P., Nyakaana, S., Douglas-Hamilton, I., & Siegismund, H. 2009. Where sociality and relatedness diverge: the genetic basis for hierarchical social structure in African elephants. Proceedings of the Royal Society: Biological Sciences 276:3513-3521 PDF

Cerling, T.E., Wittemyer, G., Ehleringer, J.R., Remien, C.H., Douglas-Hamilton I. 2009. History of Animals using Isotope Records (HAIR): A 6-year dietary history of one family of African elephants. Proceedings of the National Academy of Sciences 106:8093-8100 PDF


Wittemyer, G., Polansky, L., Douglas-Hamilton, I. and Getz, W.M. 2008. Disentangling the effects of forage, social rank and risk on movement autocorrelation of elephants using Fourier and wavelet analysis. Proceedings of the National Academy of Sciences 105:19108-19113 PDF [Supplementary Material]

Wittemyer, G.*, Brashares, J.S.*, Elsen, P., Bean, W.T. and Burton, A.C.O. 2008. Accelerated human population growth at protected area edges. Science 321:123-126 PDF [Supplementary Material]

Okello, J.B.A.*, Wittemyer, G.*, Rasmussen, H.B., Arctander, P., Nyakaana, S., Douglas-Hamilton, I., & Siegismund, H. 2008. Effective population size dynamics reveal impacts of historic climatic events and recent anthropogenic pressure in African elephants. Molecular Ecology 17:3788-3799 PDF [Supplementary Material]

Okello, J.B.A., Masembe, C., Rasmussen, H.B., Wittemyer, G., Omondi, P., Kahindi, O., Douglas-Hamilton, I., Arctander, P., Nyakaana, S., & Siegismund, H. 2008. Population genetic structure of elephants in Kenya. Journal of Heredity 99:443-452 PDF

Rasmussen, H.B., Okello, J.B.A., Wittemyer, G., Siegismund, H., Vollrath, F., & Douglas-Hamilton, I. 2008. Age and tactic related paternity success in male African elephants. Behavioral Ecology 19: 9-15 PDF


Wittemyer, G., Getz, W.M., Vollrath, F. & Douglas-Hamilton, I. 2007. Social dominance, seasonal movements, and spatial segregation in the African elephant: A contribution to conservation behavior. Behavioral Ecology and Sociobiology 61:1919-1931 PDF [Supplementary Material]

Wittemyer, G., Ganswindt, A., and Hodges, K. 2007. The impact of ecological variability on the reproductive endocrinology of wild female African elephants. Hormones and Behavior 51:346-354 PDF [Correction PDF]

Wittemyer, G.*, Rasmussen, H. B.*, & Douglas-Hamilton, I. 2007. The influence of primary productivity variability on female elephant reproductive strategies. Ecography 30:42-50 PDF

Wittemyer, G. and Getz, W.M. 2007. Testing socio-ecological predictions of dominance relationships in a scramble competitor.  Animal Behaviour 73:671-681 PDF


Wittemyer, G. and Getz, W.M. 2006. A Likely Ranking Interpolation for Resolving Optimal Dominance Orders in Systems with Unknown Relationships. Behaviour 143:909-930 PDF

Rasmussen, H. B.*, Wittemyer, G.*, & Douglas-Hamilton, I. 2006. Predicting time specific changes in demographic processes using remote sensing data. Journal of Applied Ecology 43:366-376 PDF

Cerling, T.E., Wittemyer, G., Rasmussen, H., Vollrath, F., Cerling, C.E., Robinson, T.J. & Douglas-Hamilton, I. 2006. Stable isotopes in elephant hair documents migration patterns and diet changes. Proceedings of the National Academy of Sciences 103:371-373  PDF

Douglas-Hamilton, I., Bhalla, S., Wittemyer, G., & Vollrath, F. 2006. Behavioural reactions of elephants towards a dying and deceased matriarch.  Applied Animal Behaviour Science 100:87-102 PDF


Wittemyer, G., Douglas-Hamilton, I., & Getz, W.M. 2005. The socio-ecology of   elephants: Analysis of the processes creating multileveled societies. Animal Behaviour 69:1357-1371  PDF

Wittemyer, G., Daballen, D., Rasmussen, H., Kahindi, O., Douglas-Hamilton, I. 2005. Demographic status of elephants in the Samburu and Buffalo Springs National Reserves, Kenya. African Journal of Ecology 43:44-47  PDF

Okello, J.B.A., Wittemyer, G., Rasmussen, H.B., Douglas-Hamilton, I., Nyakanna, S., Arctander, P., & Siegismund, H. 2005. Noninvasive microsatellite genotyping and Mendelian inheritance in African savanna elephants. Journal of Heredity 96:679-687  PDF

Rasmussen, H. B., Wittemyer, G., & Douglas-Hamilton, I. 2005. Estimating age of immobilized elephants from teeth impressions using dental silicon. African Journal of Ecology 43:215-219  PDF


Rasmussen, L.E.L. & Wittemyer, G. 2002.  Chemosignaling of musth by individual wild African elephants, (Loxodonta africana): implications for conservation and management. Proceedings of the Royal Society: Biological Sciences 269:853-860  PDF


Wittemyer, G. 2001.  The elephant population of Samburu and Buffalo Springs National Reserves, Kenya.  African Journal of Ecology 39:357-365  PDF

*Joint first authors