In Press

      Lynch, E.; Angeloni, L.; Fristrup, K.; Joyce, D., Wittemyer, G.  In press. The use of on-animal acoustical recording devices for studying animal behaviour. Ecology and Evolution

      Noa Pinter-Wollman, N., Hobson, E. A., Smith, J. E., Edelman, A. J., Shizuka, D., Waters, J. S., de Silva, S., Prager, S. D., Sasaki, T., Wittemyer, G., Fewell, J., McDonald, D. B. In press. The dynamics of animal social networks: analytical, conceptual, and theoretical advances. Behavioral Ecology

      Uno, K., Fisher, D., Wittemyer, G., Douglas-Hamilton, I., Cerling, T.E. In press. Bomb-curve radiocarbon measurements of recent biologic tissues and applications to stable isotope ecology and paleoecology, and to wildlife forensics. Proceedings of the National Academy of Sciences

      Gessner, J., Buchwald, R., Wittemyer, G. In press. Assessing species occurrence and species specific use patterns of bais in Central Africa with camera traps. African Journal of Ecology

      Northrup, J., Hooten, M. B., Anderson, C. R., Wittemyer, G. In press. Practical guidance on availability sampling for resource selection functions. Ecology


      Maisels, F., et al. 2013. Devastating decline of forest elephants in Central Africa.  PLoS One 8(3): e59469

      Wall, J., Wittemyer, G., Klinkenberg, B. LeMay, V., and Douglas-Hamilton, I. 2013. Characterizing properties and drivers of long distance movements by elephants (Loxodonta africana) in the Gourma, Mali. Biological Conservation 157:60-68

      Northrup, J. and Wittemyer, G. 2013. Characterizing the impacts of emerging energy development on wildlife, with an eye towards mitigation. Ecology Letters 16:112-125

      Wittemyer, G., Daballen, D., and Douglas-Hamilton, I. 2013. Comparative demography of an at risk African elephant population. PLoS One 8:e53726


      de Silva, S. and Wittemyer, G. 2012. A comparison of social organization in Asian elephants and African savannah elephants. International Journal of Primatology 33:1125-1141.


       Wittemyer, G., Daballen, D., and Douglas-Hamilton, I. 2011. Rising ivory prices threaten elephants. Nature 476:282-283.

      Bouché, P., Douglas-Hamilton, I., Wittemyer, G., Nianogo, A.J., Doucet, J., Lejeune, P., and Vermeulen, C. 2011. Will elephants soon disappear from West African savannahs? PLoS One 6:e20619

      Boetiger, A*., Wittemyer, G.*, Starfield, R., Volrath, F., Douglas-Hamilton, I., and Getz, W.M.. 2011. Inferring ecological and behavioral drivers of African elephant movement using a linear filtering approach. Ecology 92:1648-1657.

      Wittemyer, G. 2011. Effects of economic downturns on mortality of wild African elephants. Conservation Biology 25:1002-1009.

      Polansky, L.* and Wittemyer, G.* 2011. A framework for understanding the architecture of collective movements using pair-wise analyses of animal movement data. Journal of the Royal Society Interface 8:323-333.


      Polansky, L., Wittemyer, G., Cross, P.C., Tambling, C.J. & Getz, W.M. 2010. Fourier and wavelet analyses applied to animal movement time series. Ecology 91:1506-1518.

      Wasser, S., et al. 2010. Elephants, Ivory, and Trade. Science 327:1331-1332

      Kahindi, O., Wittemyer, G., King, J., Ihwagi, F., Omondi, P., & Douglas-Hamilton, I. 2010. Employing participatory surveys to monitor the illegal killing of elephants across diverse land uses in Laikipia-Samburu, Kenya. African Journal of Ecology 48:972-983


      Wittemyer, G., Cerling, T.E., & Douglas-Hamilton I. 2009. Establishing longitudinal diet chronologies from isotopic profiles in serially collected hairs. Chemical Geology 267:3-11 PDF

      Wittemyer, G., Okello, J.B.A., Rasmussen, H.B., Arctander, P., Nyakaana, S., Douglas-Hamilton, I., & Siegismund, H. 2009. Where sociality and relatedness diverge: the genetic basis for hierarchical social structure in African elephants. Proceedings of the Royal Society: Biological Sciences 276:3513-3521 PDF

      Cerling, T.E., Wittemyer, G., Ehleringer, J.R., Remien, C.H., Douglas-Hamilton I. 2009. History of Animals using Isotope Records (HAIR): A 6-year dietary history of one family of African elephants. Proceedings of the National Academy of Sciences 106:8093-8100 PDF


      Wittemyer, G., Polansky, L., Douglas-Hamilton, I. and Getz, W.M. 2008. Disentangling the effects of forage, social rank and risk on movement autocorrelation of elephants using Fourier and wavelet analysis. Proceedings of the National Academy of Sciences 105:19108-19113 PDF [Supplementary Material]

      Wittemyer, G.*, Brashares, J.S.*, Elsen, P., Bean, W.T. and Burton, A.C.O. 2008. Accelerated human population growth at protected area edges. Science 321:123-126 PDF [Supplementary Material]

      Okello, J.B.A.*, Wittemyer, G.*, Rasmussen, H.B., Arctander, P., Nyakaana, S., Douglas-Hamilton, I., & Siegismund, H. 2008. Effective population size dynamics reveal impacts of historic climatic events and recent anthropogenic pressure in African elephants. Molecular Ecology 17:3788-3799 PDF [Supplementary Material]

      Okello, J.B.A., Masembe, C., Rasmussen, H.B., Wittemyer, G., Omondi, P., Kahindi, O., Douglas-Hamilton, I., Arctander, P., Nyakaana, S., & Siegismund, H. 2008. Population genetic structure of elephants in Kenya. Journal of Heredity 99:443-452 PDF

      Rasmussen, H.B., Okello, J.B.A., Wittemyer, G., Siegismund, H., Vollrath, F., & Douglas-Hamilton, I. 2008. Age and tactic related paternity success in male African elephants. Behavioral Ecology 19: 9-15 PDF


      Wittemyer, G., Getz, W.M., Vollrath, F. & Douglas-Hamilton, I. 2007. Social dominance, seasonal movements, and spatial segregation in the African elephant: A contribution to conservation behavior. Behavioral Ecology and Sociobiology 61:1919-1931 PDF [Supplementary Material]

      Wittemyer, G., Ganswindt, A., and Hodges, K. 2007. The impact of ecological variability on the reproductive endocrinology of wild female African elephants. Hormones and Behavior 51:346-354 PDF [Correction PDF]

      Wittemyer, G.*, Rasmussen, H. B.*, & Douglas-Hamilton, I. 2007. The influence of primary productivity variability on female elephant reproductive strategies. Ecography 30:42-50 PDF

      Wittemyer, G. and Getz, W.M. 2007. Testing socio-ecological predictions of dominance relationships in a scramble competitor.  Animal Behaviour 73:671-681 PDF


      Wittemyer, G. and Getz, W.M. 2006. A Likely Ranking Interpolation for Resolving Optimal Dominance Orders in Systems with Unknown Relationships. Behaviour 143:909-930 PDF

      Rasmussen, H. B.*, Wittemyer, G.*, & Douglas-Hamilton, I. 2006. Predicting time specific changes in demographic processes using remote sensing data. Journal of Applied Ecology 43:366-376 PDF

      Cerling, T.E., Wittemyer, G., Rasmussen, H., Vollrath, F., Cerling, C.E., Robinson, T.J. & Douglas-Hamilton, I. 2006. Stable isotopes in elephant hair documents migration patterns and diet changes. Proceedings of the National Academy of Sciences 103:371-373  PDF

      Douglas-Hamilton, I., Bhalla, S., Wittemyer, G., & Vollrath, F. 2006. Behavioural reactions of elephants towards a dying and deceased matriarch.  Applied Animal Behaviour Science 100:87-102 PDF


      Wittemyer, G., Douglas-Hamilton, I., & Getz, W.M. 2005. The socio-ecology of   elephants: Analysis of the processes creating multileveled societies. Animal Behaviour 69:1357-1371  PDF

      Wittemyer, G., Daballen, D., Rasmussen, H., Kahindi, O., Douglas-Hamilton, I. 2005. Demographic status of elephants in the Samburu and Buffalo Springs National Reserves, Kenya. African Journal of Ecology 43:44-47  PDF

      Okello, J.B.A., Wittemyer, G., Rasmussen, H.B., Douglas-Hamilton, I., Nyakanna, S., Arctander, P., & Siegismund, H. 2005. Noninvasive microsatellite genotyping and Mendelian inheritance in African savanna elephants. Journal of Heredity 96:679-687  PDF

      Rasmussen, H. B., Wittemyer, G., & Douglas-Hamilton, I. 2005. Estimating age of immobilized elephants from teeth impressions using dental silicon. African Journal of Ecology 43:215-219  PDF


      Rasmussen, L.E.L. & Wittemyer, G. 2002.  Chemosignaling of musth by individual wild African elephants, (Loxodonta africana): implications for conservation and management. Proceedings of the Royal Society: Biological Sciences 269:853-860  PDF


      Wittemyer, G. 2001.  The elephant population of Samburu and Buffalo Springs National Reserves, Kenya.  African Journal of Ecology 39:357-365  PDF

      *Joint first authors