Training and Awards

  • BA Biology 1973 California State College, Sonoma
  • MA Biology 1975 California State College Sonoma – with Dr. Colin Hermans (Interstitial fauna)
  • PhD Zoology 1978 UC Berkeley – with Dr. Richard Eakin (Invertebrate photoreceptors and ciliary locomotion)
  • PostDoc UC Riverside – with Dr. Prudence Talbot (Mechanics of mammalian ovulation)
  • Occidental College 1980-present. Department Chair 1988-92; 1999-2001.
  • Loftsgordon Award for Excellence in Teaching 1984; Linda and Tod White Teaching Prize 2009
  • 1981-2008 Director of Summer Oceanology Program for select high school seniors; www.oxy.edu/oxy/marinebio/summer
  • 1996-2003 Director of the research vessel, R/V Vantuna.
  • 1996-2008 Board of Directors for Southern California Marine Institute (SCMI); www.scmi.us
  • 2003-2007 Environmental Education Advisory Committee for State of California Department of Education; www.creec.org
  • 2011-2014 Educational Council of the Society for Integrative and Comparative Biology

Membership in Professional Organizations

  • American Association for the Advancement of Science (AAAS)
  • American Society for Cell Biology (ASCB)
  • Society for Integrative and Comparative Biology (SICB)
  • The International Society of Developmental and Comparative Immunology (DCI)
  • The Crustacean Society (TCS)
  • Electron Microscopy Society of America (EMSA)
  • World Aquaculture Society (WAS)


  • Bio 115 Introduction to Zoology
  • Bio 323 Histology
  • Bio 344 Invertebrate Morphology and Physiology