Sharks, Skates, and Rays                                                     By: Jef Van Hoof

Chondrichthyes are jawed fish with paired fins, paired nares, scales, two-chambered hearts, and skeletons made of cartilage rather than bone. They are divided into two subclasses: Elasmobranchii (sharks, rays and skates) and Holocephali (chimaera, sometimes called ghost sharks, which are sometimes separated into their own class).

Sharks, skates, rays, and even stranger fish make up the Chondrichthyes, or "cartilaginous fish." First appearing on Earth almost 450 million years ago, cartilaginous fish today include both fearsome predators and harmless mollusc-eaters (harmless, that is, unless you are a mollusc). A number of shark and ray species are fished, commercially or for sport.

Members of the Chondrichthyes all lack true bone and have a skeleton made of cartilage (the flexible material you can feel in your nose and ears). Only their teeth, and sometimes their vertebrae, are calcified; this calcified cartilage has a different structure from that of true bone. Thus, preservation of the whole body of a cartilaginous fish only takes place under special conditions. This complete fossil rhinobatoid (guitarfish -- one of the earliest rays), Rhinobatis, shown on display at the Senckenberg Museum in Frankfurt, Germany, is from the Upper Cretaceous of Haqel, Lebanon, a place that has yielded many complete fossil sharks and rays.